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PRESENT TAXONOMIC ATTRIBUTIONS 
OF THE DINOFLAGELLATE CYSTS AND 

ACRITARCHS 

TREATED BY DAVEY, DOWNIE, 

SARJEANT AND WILLIAMS (1966, 1969) 



BY 

LINDA ANN KRUKEWICH 

AND 

WILLIAM ANTONY SWITHIN SARJEANT 

(University of Saskatchewan) 



PP 25-54 



BULLETIN OF 
THE BRITISH MUSEUM (NATURAL HISTORY) 
GEOLOGY Appendix II to Supplement 3 

LONDON: 1983 



© Trustees of the British Museum (Natural History), 1983 

ISSN 0524-644X 

British Museum (Natural History) 
Cromwell Road 
London SW7 5BD 



PRESENT TAXONOMIC ATTRIBUTIONS 
OF THE DINOFLAGELLATE CYSTS AND 

ACRITARCHS 

TREATED BY DAVEY, DOWNIE, 
SARJEANT AND WILLIAMS (1966, 1969) 

By LINDA A. KRUKEWICH and WILLIAM A. S. SARJEANT 

INTRODUCTION 

The major work here reprinted marked the beginning of a new epoch in systematic 
work on fossil dinoflagellate cysts, in that it embodied not only the first attempt 
to formulate a separate morphological terminology for these microfossils but also 
the first extensive application of the ideas of Evitt (1961, 1963), that archaeopyle 
character and process shape and distribution could be used as prime criteria in 
taxonomy. 

In the seventeen years that have elapsed since its publication, there has been a 
further explosive expansion in our observation and comprehension of fossil dino- 
flagellate cysts, with a consequent expansion of terminology (see Evitt et al. 1967, 
William, Sarjeant & Kidson 1978, Sarjeant 1982a). The importance in taxonomy 
of such additional characters as the shape of process terminations and the direc- 
tion of their branches, the exact shape of the archaeopyle, the presence and 
pattern of minute plates (we would now style them "paraplates") in the mid- 
ventral region, and the exact relationships of the anterior ventral plates has come 
to be recognized (see for example Evitt 1967, Lentin & Williams 1976, Sarjeant, 
1982b). In contrast, some features - for example, the presence of an ectophragm 
- are now considered less reliable as means for the separation of taxa. The result 
has been the reassignment to other genera of many of the species mentioned in 
the 1966 paper and its first Appendix (1969). The diagnoses of many of these 
genera and species have been greatly modified, a few taxa have been rejected as 
junior synonyms and some species, then treated as acritarchs, are now recognized 
to be dinoflagellate cysts. 

To simplify understanding of the present taxonomic situation, a checklist is 
here appended in which the names of all genera and species mentioned in 1966 
and in the 1969 Appendix are given in alphabetical order. Any revisions to their 
diagnoses are noted; and the present names, where different to those cited earlier, 
are indicated. Where their taxonomic allocation remains a matter for controversy, 
the alternative ideas are noted. The species placed in 1966 into the acritarch genus 
Baltisphaeridium are included in proper alphabetical position; if these are still 
considered to be acritarchs, that fact is likewise noted. Where taxa appeared 
under different generic names in 1966 and 1969, both names are included in the 
list. A full reference section, citing all papers referred to in the list, is appended. 



28 



SECOND APPENDIX TO 



Two papers by the second author which are currently in press (Sarjeant 
1983a, b) , and may not be published until after this checklist has appeared, include 
further revisions to the taxonomy of some of these species mentioned herein. Both 
are restudies of European type material; the first deals with the Oligocene to 
Miocene taxa described by Maier (1959) and the second with the Jurassic types of 
Klement (i960). The imminent revisions are not cited here, to avoid priority 
problems; but the Maier species concerned are marked with a single asterisk (*), 
indicating that reference should also be made to Sarjeant (1983a), and the Kle- 
ment species with a double asterisk (**), suggesting reference to Sarjeant (1983b). 



CHECKLIST 



Past Name 
ACANTHOGONYAULAX Sarjeant, 1966b 



venusta (Klement, i960) Sarjeant, 1966b 

ACHOMOSPHAERA Evitt, 1963 

alcicornu (Eisenack, 1954) Davey & 

Williams, 1966a 
grallaeformis (Brosius, 1963) Davey & 

Williams, 1969 
hirundo (Eisenack, 1958) Davey & Williams, 

1969 
hyperacantha (Deflandre & Cookson, 1955) 

Davey & Williams, 1969 
neptuni (Eisenack, 1958) Davey & Williams, 

1966a 
ramulifera (Deflandre, 1937) Evitt, 1963 
ramulifera var. perforata Davey & Williams, 

1966a 
sagena Davey & Williams, 1966a 
triangulata (Gerlach, 1961) Davey & 

Williams, 1969 

ADNATOSPHAERIDIUM Williams & 

Downie, 1966c 
aemulum (Deflandre, 1938) Williams & 

Downie, 1969 
caulleryi (Deflandre, 1938) Williams & 

Downie, 1969 
filamentosum (Cookson & Eisenack, 1958) 

Williams & Downie, 1969 
multispinosum Williams & Downie, 1966c 
patulum Williams & Downie, 1966c 

vittatum Williams & Downie, 1966c 



Present Name 

ACANTHA ULAX Sarjeant, 1968 [The name 
Acanthogonyaulax was ajr horn, of a sub- 
genus of Gonyaulax proposed by Kofoid, 
191 1. The subgenus was elevated to 
generic status by Graham, 1942.] 

Acanthaulax venusta (Klement, i960) Sar- 
jeant, 1968 



Cordosphaeridiutn? hirundo (Eisenack, 1958) 
Stover & Evitt, 1978 

Spiniferites hyper acanthus (Deflandre & Cook- 
son, 1955) Cookson & Eisenack, 1974 



[Treated by Lentin & Williams, 1973, as a sub- 
species.] 



Rigaudella aemula (Deflandre, 1938) Below, 
1982 



Rigaudella fdamentosa (Cookson & Eisenack, 
1958) Below, 1982 

Thalassiphora patula (Williams & Downie, 
1966c) Stover & Evitt, 1978 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 



29 



APTEODINIUM Eisenack, 1958^ 
granulatum Eisenack, 1958 
maculatum Eisenack & Cookson, i960 

AREOLIGERA Lejeune-Carpentier, 1938; 
emend. Williams & Downie, 1966c 

coronata (O. Wetzel, 1933) Lejeune-Carpen- 
tier, 1938 

galea Maier, 1959* 

lychnea (Maier, 1959) Davey et al., 1969* 

twistringensis (Maier, 1959) Davey et al., 
1969* 

BALTISPHAERIDIUM Eisenack, 1958; 
emend. Downie & Sarjeant, 1963; emend. 
Eisenack, 1969 

armatum (Deflandre, 1937) Downie & Sar- 
jeant, 1963 

asteroideum (Maslov, 1956) Downie & Sar- 
jeant, 1963 

claviculorum (Deflandre, 1938) Downie & 
Sarjeant, 1963 

clavispinulosum Churchill & Sarjeant, 1963 
densicomatum (Maier, 1959) Gerlach, 1961* 
denticulatum (Courtville in Deflandre, 1946) 

Downie & Sarjeant, 1963 
Pdifforme (Pritchard, 1841) Downie & Sar- 
jeant, 1963 
downiei Sarjeant, i960 

echiniplax Churchill & Sarjeant, 1963 

fimbriatum (White, 1842) Sarjeant, 1959 

gilsonii (Kufferath, 1950) Downie & Sar- 
jeant, 1963 

horridum (Deflandre, 1937) Downie & Sar- 
jeant, 1963 

huguonioti (Valensi, 1955) Downie & Sar- 
jeant, 1963 

intermedium (O. Wetzel, 1933) Downie & 
Sarjeant, 1963 

longofilum (Maier, 1959) Downie & Sarjeant, 
1963* 

malleo/erum (White, 1842) Downie & Sar- 
jeant, 1963 

mariannae (Philippot, 1949) Downie & Sar- 
jeant, 1963 

pannifor me Gerlach, 1961 

pattei (Valensi, 1948) Sarjeant, i960 

paucifurcatum (Cookson & Eisenack, 1961) 

Downie & Sarjeant, 1965 
pectiniforme Gerlach, 1961 



Cleistosphaeridium armatum (Deflandre, 1937) 
Davey, 1969a 
Regarded as an acritarch. 

Solisphaeridium claviculorum Deflandre, 1938; 
emend. Sarjeant, 1968 
Regarded as an acritarch. 
Regarded as an acritarch. 

Regarded as an acritarch. 



Acanthaulax? downiei (Sarjeant, i960) Sar- 
jeant, 1976 
Regarded as an acritarch. 
Regarded as an acritarch. 
Regarded as an acritarch. 

Regarded as an acritarch. 

Cleistosphaeridium huguoniotii (Valensi, 1955) 

Davey, 1969a 
Hystrichosphaeridium intermedium (O. Wetzel, 

1933) Deflandre, 1937 
Regarded as an acritarch. 

Regarded as an acritarch. 

Achomosphaera mariannae (Philippot, 1949) 

Stover & Evitt, 1978 
Cleistosphaeridium panniforme (Gerlach, 1961) 

Stover & Evitt, 1978 
Lanterna? pattei (Valensi, 1948) Brideaux & 

Fisher, 1976 
Regarded as an acritarch. 

Areosphaeridium? pectiniforme (Gerlach, 1961) 
Stover & Evitt, 1978 



30 



SECOND APPENDIX TO 



BALTISPHAERIDIUM continued 

plicatutn (Maier, 1959) Downie & Sarjeant, 

1963* 
Ppolyceratum Takahashi, 1964 
polyozon Brosius, 1963 

quaternarium Churchill & Sarjeant, 1963 

saturnium (Maier, 1959) Downie & Sarjeant, 
1963* 

seminudum (W. Wetzel, 1952) Downie & 
Sarjeant, 1963 

spiculatum (White, 1842) Downie & Sar- 
jeant, 1963 

stimuliferum (Deflandre, 1938) Sarjeant, 
i960 

sylheti (Baksi, 1962) Downie & Sarjeant, 

1963 
telmaticum Churchill & Sarjeant, 1963 
tinglewoodense Churchill & Sarjeant, 1963 
varispinosum Sarjeant, 1959 



whitei (Deflandre & Courteville, 
Downie & Sarjeant, 1963 



1939) 



BELODINIUM Cookson & Eisenack, 1960b; 
emend. Dodekova, 1975 
dysculum Cookson & Eisenack, 1960b 



Regarded as an acritarch. 
Cleistosphaeridium? polyozon (Brosius, 1963) 
Stover & Evitt, 1978 
Regarded as an acritarch. 



Regarded as an acritarch. 

Regarded as an acritarch. 

Solisphaeridium stimuliferum (Deflandre, 
1938) Staplin, Jansonius & Pocock, 1965 
Regarded as an acritarch. 
Hystrichosphaeridium sylheti Baksi, 1962 

Regarded as an acritarch. 
Regarded as an acritarch. 
SentusidiniumP varispinosum (Sarjeant, 1959) 

Sarjeant & Stover, 1978 
Cometodinium? whitei (Deflandre & Courte- 
ville, 1939) Stover & Evitt, 1978 






BROOMEA Cookson & Eisenack, 1958; 
emend. Lentin & Williams, 1976 
Plongicornuta Alberti, 196 1 

ramosa Cookson & Eisenack, 1958 

CALLAIOSPHAERIDIUM Davey & 
Williams, 1966b; emend. Duxbury, 1980; 
emend. Below, 1981 
asymmetricum (Deflandre & Courtville, 
1939) Davey & Williams, 1966b 

CANNOSPHAEROPSIS O. Wetzel, 1933; 
emend. Williams & Downie, 1966c; 
emend. May, 1980; emend. Duxbury, 1980 
reticulensis Pastiels, 1948 

utinensis O. Wetzel, 1933; emend. Duxbury, 
1980 

CA RPODINIUM Cookson & Eisenack, 1962; 
emend. Lefhngwell & Morgan, 1977 
granulatum Cookson & Eisenack, 1962; 
emend. Lefhngwell & Morgan, 1977 



Imbatodinium longicornutum (Alberti, 1961) 
Dorhofer & Davies, 1980 



Adnatosphaeridium reticulense (Pastiels, 1948) 
De Coninck, 1969 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 



3i 



CLEISTOSPHAERIDIUM Davey et al., 

1966 
ancoriferum (Cookson & Eisenack, 1960a) 

Davey et al., 1966; emend. Cookson & 

Eisenack, 1968 
ashdodense Rossignol, 1962 

danicum (W. Wetzel, 1952) Davey et al., 

1969 
disjunctum Davey et al., 1966 

diver si spino sum Davey et al., 1966 
echinoides (Maier, 1959) Davey et al., 1969* 
ehrenbergii (Deflandre, 1947) Davey et al., 

1969 
Pflexuosum Davey et al., 1966 
heteracanthum (Deflandre & Cookson, 1955) 

Davey et al., 1966 

israelianum (Rossignol, 1962) Davey et al., 

1966 
leve (Maier, 1959) Davey et al., 1969* 
lumectum (Sarjeant, 1960a) Davey et al., 

1969 
machaerophorum (Deflandre & Cookson, 

1955) Davey et al., 1966 
multifurcatum (Deflandre, 1937) Davey et 

al., 1969 
Poligacanthum (W. Wetzel, 1952) Davey et 

al., 1969 
pectiniforme (Gerlach, 1961) Davey et al., 

1969 
pilosum (Ehrenberg, 1843) Davey et al., 1966 



polypes (Cookson & Eisenack, 1962) Davey, 

1969a 
polytrichum (Valensi, 1947) Davey et al., 

1969 
spiralisetum (De Wit, 1943) Davey et al., 

1969 
tiara (Klumpp, 1953) Davey et al., 1969 
tribuliferum (Sarjeant, 1962b) Davey et al., 

1969 

COMETODINIUM Deflandre & Courteville, 

1939 
obscurum Deflandre & Courteville, 1939 

CORDOSPHAERIDIUM Eisenack, 1963b; 
emend. Davey, 1969c; emend. Sarjeant, 
1981 
cantharellum (Brosius, 1963) Gocht 1969 



[Wall, 1967 considered this species to be a.jr 
syn. of Lingulodinium machaerophorum] 



[Reid, 1974, considered this species to be a 
jr syn. of Lingulodinium machaerophorum] 



Heterosphaeridium? heteracanthum (Deflandre 
& Cookson, 1955) Eisenack & Kjellstrom, 
1971 

Operculodinium israelianum (Rossignol, 1962) 
Wall, 1967 



Lingulodinium machaerophorum (Deflandre & 
Cookson, 1955) Wall, 1967 



A reosphaeridium? pectiniforme (Gerlach, 1961) 
Stover & Evitt, 1978 

Sentusidinium pilosum (Ehrenberg, 1843) Sar- 
jeant & Stover, 1978; emend. Erkmen & 
Sarjeant, 1980 

Bacchidinium polypes (Cookson & Eisenack, 
1962) Davey, 1979 



Holotype lost; exact character and sys- 
tematic status problematical. 



Tityrosphaeridium cantharellum (Brosius, 
1963) Sarjeant, 1981 



32 



SECOND APPENDIX TO 



CORDOSPHAERIDIUM continued 
Pcracenospinosum Davey & Williams, 1966b 

diktyoplokus Klumpp, 1953 

diktyoplokus subsp. latum Klump, 1953 

divergens (Eisenack, 1954) Eisenack, 1963b 

eoinodes (Eisenack, 1954) Eisenack, 1963b 

Perectum (Manum & Cookson, 1964) Davey 

& Williams, 1969 
exilimurum Davey & Williams, 1966b 

Pfasciatum Davey & Williams, 1966b 

fibrospinosum Davey & Williams, 1966b 

ftoripes (Deflandre & Cookson, 1955) Eisen- 
ack, 1963b 

floripes subsp. breviradiatum (Cookson & 
Eisenack, 1961b) Eisenack, 1963b 

gracile (Eisenack, 1954) emend. Davey & 
Williams, 1966b 

latispinosum Davey & Williams, 1966b 



microtriainum (Klump, 1953) Eisenack, 

1963b 
multispinosum Davey & Williams, 1966b 

CTENIDODINIUM Deflandre, 1938; emend. 
Sarjeant, 1966b; emend. Sarjeant, 1975 
ornatum (Eisenack, 1935) Deflandre, 1938 
tenellum Deflandre, 1938 

CYCLONEPHELIUM Deflandre & Cookson, 
1955; emend. Williams & Downie, 1966c; 
emend. Ioannides et al., 1976; emend. Sar- 
jeant & Stover, 1978; emend. Stover & 
Evitt, 1978 
compactum Deflandre & Cookson, 1955 
divaricatum Williams & Downie, 1966c 

exuberans Deflandre & Cookson, 1955 

ordinatum Williams & Downie, 1966c 

pastielsii Deflandre & Cookson, 1955 



Achomosphaera? cracenospinosa (Davey & 
Williams, 1966b) Sarjeant, 1981 

Areosphaeridum diktyoplokus (Klump, 1953) 
Eaton, 1971 

Areosphaeridium diktyoplokus subsp. latum 
(Klumpp, 1953) Lentin & Williams, 1973 

Operculodinium divergens (Eisenack, 1954) 
Stover & Evitt, 1978 

Kleithriasphaeridium eoinodes (Eisenack, 
1958) Davey, 1974 

Cleistospkaeridium? erectum (Manum & Cook- 
son, 1964) Stover & Evitt, 1978 

Hystrichosphaerina? exilimura (Davey & 
Williams, 1966b) Sarjeant, 1981 

Kleithriasphaeridium fasciatum (Davey & 
Williams, 1966b) Davey, 1974 

Emmetrocysta? fibrospinosa (Davey & 
Williams, 1966b) Sarjeant, 1981 

Homotryblium floripes (Deflandre & Cookson, 
1955) Stover, 1975 

Homotryblium floripes subsp. breviradiatum 
(Cookson & Eisenack, 1961b) Lentin & 
Williams, 1977 

Tityrosphaeridium gracilis (Eisenack, 1954 
emend. Davey & Williams, 1966b) Sar- 
jeant, 1981 

Titryosphaeridium? latispinosum (Davey & 
Williams, 1966b) Sarjeant, 1981 [Placed 
earlier into A chilleodinium by Bujak etal., 
1980; this may prove to be a more correct 
assignment.] 

Achomosphaera microtriaina (Klump, 1953) 
emend. Sarjeant, 1981 

Amphorosphaeridum? multispinosum (Davey 
& Williams, 1966b) Sarjeant, 1981 



Glaphyrocysta divaricata (Williams & Downie, 
1966c) Stover & Evitt, 1978 

Glaphyrocysta exuberans (Deflandre & Cook- 
son, 1955) Stover & Evitt, 1978 

Glaphyrocysta ordinata (Williams & Downie, 
1966c) Stover & Evitt, 1978 

Glaphyrocysta pastielsii (Deflandre & Cookson, 
1955) Stover & Evitt, 1978 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 



33 



CYMATIOSPHAERA O. Wetzel, 1933; 
emend. Deflandre, 1954 
membranacea (Philippot, 1949) Davey & 
Williams, 1969 

DEFLANDRE A Eisenack, 1938; emend. 

Williams & Downie, 1966c; emend. Lentin 

& Williams, 1976 
denticulata Alberti, 1959 
oebisfeldensis Alberti, 1959 
phosphoritica Eisenack, 1938 
phosphoritica subsp. australis Cookson & 

Eisenack, 1961a 
phosphoritica subsp. phosphoritica Eisenack, 

1938 
wardenensis Williams & Downie, 1966c 

DICHADOGONYAULAX Sarjeant, 1966a 

culmula (Norris, 1965) Loeblich & Loeblich, 

1968 
pannea (Norris, 1965) Sarjeant, 1969 

schizoblatum (Norris, 1965) Sarjeant, 1969 

DINGODINIUM Cookson & Eisenack, 1958 
? albertii Sarjeant, 1966c 
jurassicum Cookson & Eisenack, 1958 

DIPHYES Cookson, 1965; emend. Davey & 

Williams, 1966b 
colligerum (Deflandre & Cookson, 1955) 

Cookson, 1965 
Pmonstruosum (Tasch, 1964) Davey & 

Williams, 1969 

DOIDYX Sarjeant, 1966b 



anaphrissa Sarjeant, 1966c 

DUOSPHAERIDIUM Davey & Williams, 
1966b 
nudum (Cookson, 1965) Loeblich & Loe- 
blich, 1968 

EISENACKIA Deflandre & Cookson, 1955; 
emend. Sarjeant, 1966b; emend. McLean, 
1973; emend. Stover & Evitt, 1978 
crassitabulata Deflandre & Cookson, 1955; 
emend. McLean, 1973 



Regarded as an acritarch. 



Ceratiopsis wardenensis (Williams & Downie, 
1966c) Bujak et al., 1980 

[Considered by Lentin & Williams, 1973 to 

be ajr syn. of Ctenidodinium] 
[Transferred to Ctenidodinium by Lentin & 

Williams, 1973.] 
[Transferred to Ctenidodinium by Lentin & 

Williams, 1973.] 
[Transferred to Ctenidodinium by Lentin & 

Williams, 1973] 



Coronifera monstruosa (Tasch, 1964) Stover & 
Evitt, 1978 

Shown by Sarjeant & Stover, 1978 to be a.jr 
syn. of APTEA Eisenack, 1958; emend. 
Davey & Verdier, 1974; emend. D6rhofer& 
Davies, 1980 
Aptea anaphrissa (Sarjeant, 1966c) Stover & 
Sarjeant, 1978 



34 



SECOND APPENDIX TO 



EXOCHOSPHAERIDIUM Davey et at., 

1966; emend. Davey, 1969c 
palmatum (Deflandre & Courtville, 1939) 

Davey et at., 1969 
phragmites Davey et at., 1966 
pseudhystrichodinium (Deflandre, 1937) 

Davey et at., 1969 
striolatum (Deflandre, 1937) Davey, 1969a 

F ROME A Cookson & Eisenack, 1958 
amphora Cookson & Eisenack, 1958 

GARDODINIUM Alberti, 1961 
eisenackii Alberti, 1961 



GLYPHANODINIUM Drugg, 1964 
facetum Drugg, 1964 

GONYAULACYSTA Deflandre, 1964; 
emend. Sarjeant, 1969; emend. Stover & 
Evitt, 1978; emend. Sarjeant, 1982a 
Paceras (Eisenack, 1958) Sarjeant, 1966b 

aculeata (Klement, i960) Sarjeant, 1969 

aichmetes Sarjeant, 1966b 

amabilis Deflandre, 1939 

ambigua (Deflandre, 1939) Sarjeant, 1969 

apionis (Cookson & Eisenack, 1958) Sar- 
jeant, 1969 

aptiana (Deflandre, 1935) Sarjeant, 1966b 

axicerastes Sarjeant, 1966b 

cassidata (Eisenack & Cookson, i960) Sar- 
jeant, 1966b 

cladophora (Deflandre, 1938) Sarjeant, 
1966b 

clathrata (Cookson & Eisenack, 1960b) Sar- 
jeant, 1966b 

confusa (Vozzhenikova, 1967) Sarjeant, 
1969 

Pcornigera (Valensi, 1953) Sarjeant, 1966b 

crassicornuta (Klement, i960) Sarjeant, 

1966b 
cretacea (Neale & Sarjeant, 1962) Sarjeant^ 

1969 
crispa (W. Wetzel, 1967) Sarjeant, 1969 

dangeardii Sarjeant, 1968 



Coronifera striolata (Deflandre, 1937) Stover 
& Evitt, 1978 



[Davey, 1974, considered this species to be a.jr 
syn. of Gardodinium trabeculosum (Gocht, 
1959) Alberti, 1961.] 



Acanthaulax? aceras (Eisenack, 1958) Stover 
& Evitt, 1978 

Acanthaulax aculeata (Klement, i960) Stover 
& Evitt, 1978 

Millioudodinium aichmetes (Sarjeant, 1966b) 
Stover & Evitt, 1978 

Leptodinium? amabile (Deflandre, 1939) Sar- 
jeant, 1969 

Rhynchodiniopsis ambigua (Deflandre, 1939) 
Sarjeant, 1982a 

Millioudodinium apionis (Cookson & Eisen- 
ack, 1958) Stover & Evitt, 1978 

Rhynchodiniopsis aptiana Deflandre, 1935 



Rhynchodiniopsis cladophora (Deflandre, 1938) 

Below, 1 98 1 
Leptodinium clathratum (Cookson & Eisenack, 

1960b) Sarjeant, 1969 
Rhynchodiniopsis? confusa (Vozzhennikova, 

1967) Sarjeant, 1982a 
Hystrichogonyaulax cornigera (Valensi, 1953) 

Sarjeant, 1969 
Rhynchodiniopsis crassicornuta (Klement, 

i960) Below, 1981 



Hystrichogonyaulax crispa (W. Wetzel, 1967) 

Sarjeant, 1980b 
Tubotuberella dangeardii (Sarjeant, 1968) 

Stover & Evitt, 1978 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 



35 



GONYAULACYSTA continued 

diaphanis (Cookson & Eisenack, 1958) Sar- 

jeant, 1969 
dictyophora (Deflandre, 1938) Sarjeant, 1967 

edwardsii (Cookson & Eisenack, 1958) 

Clarke & Verdier, 1967 
eisenackii (Deflandre, 1938) Dodekova, 

1967; emend. Sarjeant, 1982a 
episoma Sarjeant, 1966b 

eumorpha (Cookson & Eisenack, 1960b) Sar- 
jeant, 1966b 
exilicristata Davey, 1969a 

fetchamensis Sarjeant, 1966b 

freakei (Sarjeant, 1963) Sarjeant, 1966a 

giuseppei (Morgenroth, 1966) Sarjeant, 1969 

gongylos Sarjeant, 1966b 

gottisii Dupin, 1968 

granulata (Klement, i960) Sarjeant, 1969** 

granuligera (Klement, i960) Sarjeant, 

1969** 
hadra Sarjeant, 1966b 

helicoidea (Eisenack & Cookson, i960) Sar- 
jeant, 1966b 
hyaloderma (Deflandre, 1939) Sarjeant, 1967 

hyalodermopsis (Cookson & Eisenack, 1958) 
Sarjeant, 1969 

jurassica (Deflandre, 1938) Norris & Sar- 
jeant, 1965; emend. Sarjeant, 1982a 

kostromiensis (Vozzhennikova, 1967) Sar- 
jeant, 1969 

longicornis (Downie, 1957) Sarjeant, 1969 

?mamillifera (Deflandre, 1939) Sarjeant, 
1969 

margaritifera (Cookson & Eisenack, 1960a) 
Sarjeant, 1966b 

microceras (Eisenack, 1958) Clarke & Ver- 
dier, 1967 

millioudi (Sarjeant, 1963) Sarjeant, 1966b 

tnonacantha (Deflandre, 1935) Sarjeant, 
1967 



Cribroperidinium diaphanis (Cokson & Eisen- 
ack, 1958) Stover & Evitt, 1978 

Aldorfia dictyophora (Deflandre, 1938) Stover 
& Evitt, 1978 

Cribroperidinium edwardsii (Cookson & 
Eisenack, 1958) Davey, 1969a 



Rhynchodiniopsis episoma (Sarjeant, 1966b) 
Sarjeant, 1982a 

Leptodinium eumorphum (Cookson & Eisen- 
ack, 1960b) Sarjeant, 1969 

Cribroperidinium exilicristatum (Davey, 
1969a) Stover & Evitt, 1978 

Millioudodinium fetchamense (Sarjeant, 
1966b) Stover & Evitt, 1978 

Leptodinium freakei (Sarjeant, 1963) Sarjeant, 
1969 

Rhynchodiniopsis? giuseppeii (Morgenroth, 
1966) Sarjeant, 1982a 

Rhynchodiniopsis gongylos (Sarjeant, 1966b) 
Sarjeant, 1982a 

Millioudodinium? gottisii (Dupin, 1968) Stover 
& Evitt, 1978 

Rhynchodiniopsis granulata (Klement, i960) 
Sarjeant, 1982a 

Rhynchodiniopsis granuligera (Klement, i960) 
Sarjeant, 1982a 

Rhynchodiniopsis hadra (Sarjeant, 1966b) Sar- 
jeant, 1982a 



Rhynchodiniopsis? hyaloderma (Deflandre, 

1939) Sarjeant, 1982a 
Rhynchodiniopsis hyalodermopsis (Cookson & 

Eisenack, 1958) Sarjeant, 1982a 



N elchinopsis kostromiensis (Vozzhennikova, 
1967) Wiggins, 1972 [The transfer of this 
species to N elchinopsis was rejected by 
Duxbury, 1977] 

Millioudodinium longicornis (Downie, 1957) 
Sarjeant, 1982a 

Rhynchodiniopsis? mamillifera (Deflandre, 
1939) Sarjeant, 1982a 

I mpagidinium margaritiferum (Cookson & 
Eisenack, 1960a) Stover & Evitt, 1978 



Leptodinium millioudii (Sarjeant, 1963) Sar- 
jeant, 1969 

Millioudodinium monacanthum (Deflandre, 
1935) Stover & Evitt, 1978 



36 



SECOND APPENDIX TO 



GONYAULACYSTA continued 

muderongensis (Cookson & Eisenack, 1958) 

Sarjeant, 1966b 
Pnannotrix (Deflandre, 1939) Sarjeant, 1969 

Pnealei (Sarjeant, 1962b) Sarjeant, 1966b 

nuciformis (Deflandre, 1938) Sarjeant, 1968 

obscura (Lejeune-Carpentier, 1946) Sar- 
jeant, 1969 

orthoceras (Eisenack, 1958) Sarjeant, 1966b 

pachyderma (Deflandre, 1938) Sarjeant, 

1969 
palla Sarjeant, 1966b 

par orthoceras Davey, 1968 

per for ans (Cookson & Eisenack, 1958) Sar- 
jeant, 1969 

Pporosa (Lejeune-Carpentier, 1946) Sar- 
jeant, 1966b 

pyra (Drugg, 1967) Sarjeant, 1969 

sarjeantii (Vozzhennikova, 1967) Sarjeant, 
1969 

scar bur ghensis Sarjeant, 1964 

scottii (Cookson & Eisenack, 1958) Sarjeant, 
1969 

serrata (Cookson & Eisenack, 1958) Sar- 
jeant, 1969 

Ptenuiceras (Eisenack, 1958) Sarjeant, 1969 

tenuicornuta (Cookson & Eisenack, 1962) 
Sarjeant, 1969 

tenuitabulata (Gerlach, 1961) De Coninck, 
1969 

Ptransparens (Sarjeant, 1959) Sarjeant, 1969 

wetzelii (Lejeune-Carpentier, 1939) Sarjeant, 
1969; emend. Lejeune-Carpentier & Sar- 
jeant, 1981 

whitei Sarjeant, 1966b 

HELIODINIUM Alberti, 1961 



patriciae Neale & Sarjeant, 1962 
voigti Alberti, 1961 



Cribroperidinium muderongense (Cookson & 
Eisenack, 1958) Davey, 1969a 

RhynchodiniopsisP nannotrix (Deflandre, 1939) 
Sarjeant, 1982a 

HystrichogonyaulaxP nealei (Sarjeant, 1962b) 
Sarjeant, 1969 

Millioudodinium nuciformis (Deflandre, 1938) 
Sarjeant, 1982a 

Millioudodinium obscurum (Lejeune-Carpen- 
tier, 1946; emend. Lejeune-Carpentier & 
Sarjeant, 1981) Sarjeant, 1982a 

Cribroperidinium orthoceras (Eisenack, 1958) 
Davey, 1969a 

CtenidodiniumP pachydermum (Deflandre, 
1938) Gocht, 1970 

Rhynckodiniopsis palla (Sarjeant, 1966b) Sar- 
jeant, 1982a 

Millioudodinium parorthoceras (Davey, 1968) 
Stover & Evitt, 1978 

Rhynckodiniopsis perforans (Cookson & Eisen- 
ack, 1958) Sarjeant, 1982a 

Leptodinium porosum (Lejeune-Carpentier, 
1946) Sarjeant, 1969; emend. Lejeune- 
Carpentier & Sarjeant, 1981. [Conceivably 
a fossil motile dinoflagellate!] 

Cribroperidinium? pyrum (Drugg, 1967) Stover 
& Evitt, 1978 

Rhynckodiniopsis sarjeantii (Vozzhennikova, 
1967) Sarjeant, 1982a 

Millioudodinium? scottii (Cookson & Eisenack, 
1958) Stover & Evitt, 1978 

Hystrichogonyaulax serrata (Cookson & Eisen- 
ack, 1958) Stover & Evitt, 1978 

Diacanthum tenuiceras (Eisenack, 1958) Stover 
& Evitt, 1978 

Leptodinium? tenuicornutum Cookson & 
Eisenack, 1962 

Millioudodinium tenuitabulatum (Gerlach, 
1961) Stover & Evitt, 1978 



Rhynckodiniopsis whitei (Sarjeant, 1966b) Sar- 
jeant, 1982a 

[Considered by Lentin & Williams, 1975 to 
be ■a.jr syn. of Hystrichodinium Deflandre, 
1935. This judgement, and the resultant 
transfer of its two constituent species to 
Hystrichodinium, are not accepted by us.] 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 



37 



HEMICYSTODINIUM Alberti, 1961 
zoharyi (Rossignol, 1962) Wall, 1967 

HESLERTONIA Sarjeant, 1966b; emend. 
Duxbury, 1980 
heslertonensis (Neale & Sarjeant, 1962) Sar- 
jeant, 1966b 

HOMOTRYBLIUMT>avey& Williams, 1966b 
pallidum Davey & Williams, 1966b 
tenuispinosum Davey & Williams, 1966b 

HYSTRICHODINIUM Deflandre, 1935; 
emend. Clarke & Verdier, 1967 
compactum Alberti, 1961 
oligacanthum Deflandre & Cookson, 1955 
pulchrum Deflandre, 1935 
ramoides Alberti, 1961 

HYSTRICHOGONYAULAX Sarjeant, 1969 
cornigera (Valensi, 1953) Sarjeant, 1969 
nealei (Sarjeant, 1962b) Sarjeant, 1969 

HYSTRICHOKOLPOMA Klumpp, 1953; 

emend. Williams & Downie, 1966a 
cinctum Klumpp, 1953 
clavigerum (Deflandre, 1937) Williams & 

Downie, 1969 

eisenacki Williams & Downie, 1966a 



eisenacki var. turgidum Williams & Downie, 

1966a 
ferox (Deflandre, 1937) Davey, 1969a 

poculum Maier, 1959* 

rigaudae Deflandre & Cookson, 1955 

sequanaportus Deflandre & Deflandre- 

Rigaud, 1958 
unispinum Williams & Downie, 1966a 
Pxipheum (Maier, 1959) Davey & Williams, 

1969* 

HYSTRICHOSPHAERA O. Wetzel, 1933 ex 
Deflandre, 1937 
buccina Davey & Williams, 1966a 



cingulata (O.Wetzel, 1933) Deflandre, 1954 

cingulata var. reticulata Davey & Williams, 
1966a 

cornuta Gerlach, 1961 



Jr syn. of Polysphaeridium: see Bujak et al., 

1980 
Polysphaeridium zoharyi (Rossignol, 1962) 

Bujak et al., 1980 



Florentinia clavigera (Deflandre, 1937) Davey 

& Verdier, 1973; emend. Davey & Verdier, 

1976 
[Considered by Damassa, 1979, to be a jr 

syn. of Hystrichokolpoma cinctum 

Klumpp, 1953.] 
Hystrichokolpoma cinctum subsp. turgidum 

Lentin & Williams, 1973 
Florentinia ferox (Deflandre, 1937) Duxbury, 

1980 



Shown by Sarjeant, 1970 to be a jr syn. of 

Spinijerites Mantell, 1850 
Shown by Lentin & Williams, 1973, to be a. jr 

syn. of Spinijerites pseudofurcatus 

(Klumpp, 1953) Sarjeant, 1970; emend. 

Sarjeant, 1981. 
Pterodinium cingulatum (O. Wetzel, 1933) 

Below, 1981 
Pterodinium cingulatum subsp. reticulatum 

(Davey & Williams, 1966a) Lentin & 

Williams, 1981 
Spinijerites cornutus (Gerlach, 1961) Sarjeant, 

1970 



3« 



SECOND APPENDIX TO 



HYSTRICHOSPHAERA continued 

cornuta var. laevimura Davey & Williams, 

1966a 
crassimurata Davey & Williams, 1966a 

crassipellis Deflandre & Cookson, 1955 

lepioderma (Maier, 1959) Davey & Williams, 

1969* 
monolis Davey & Williams, 1966a 

perforata Davey & Williams, 1966a 

ramosa (Ehrenberg, 1838) O. Wetzel, 1933 

ramosa var. gracilis Davey & Williams, 
1966a 



ramosa var. granosa Davey & Williams, 
1966a 



ramosa var. granomembranacea Davey & 
Williams, 1966a 



ramosa var. membranacea (Rossignol, 1964) 

Davey & Williams, 1966a 
ramosa var. multibrevis Davey & Williams, 

1966a 



ramosa var. ramosa (Ehrenberg, 1838) 
Davey & Williams, 1966a 



ramosa var. reticulata Davey & Williams, 
1966a 



HYSTRICHOSPHAERIDIUM Deflandre, 
1937; emend. Davey & Williams, 1966b 
Paquitanicum Deunff, 1961 
arborispinum Davey & Williams, 1966b 
Parundum Eisenack & Cookson, i960 

bowerbankii Davey & Williams, 1966b 
claviferum (Wilkinson, 1849) Deflandre, 

1946 
Pclavigerum (Deflandre, 1937) Lejeune- 

Carpentier, 1940 

costatum Davey & Williams, 1966b 



[Treated as a subspecies by Lentin & 
Williams, 1973.] 

Spiniferites crassimuratus (Davey & Williams, 
1966a) Sarjeant, 1970 

Achomosphaera crassipellis (Deflandre & Cook- 
son, 1955) Stover & Evitt, 1978 



Spiniferites monolis (Davey & Williams, 
1966a) Sarjeant, 1970 

Spiniferites perforatus (Davey & Williams, 
1966a) Sarjeant, 1970 

Spiniferites ramosus (Ehrenberg, 1838) Man- 
tell, 1857 

Spiniferites ramosus var. gracilis (Davey & 
Williams, 1966a) Corradini, 1973 [Treated 
by Lentin & Williams, 1981, as a sub- 
species] 

Spiniferites ramosus var. granosus (Davey & 
Williams, 1966a) Corradini, 1973 [Treated 
by Lentin & Williams, 1981, as a sub- 
species] 

Spiniferites ramosus var. granomembranaceus 
(Davey & Williams, 1966a) Corradini, 
1973 [Treated by Lentin & Williams, 
1 98 1, as a subspecies] 

Spiniferites membranaceus (Rossignol, 1964) 
Sarjeant, 1970 

Spiniferites ramosus var. multibrevis (Davey 
& Williams, 1966a) Davey & Verdier, 1971 
[Treated by Lentin & Williams, 1981, as 
a subspecies] 

Spiniferites ramosus var. ramosus (Ehrenberg, 
1838) Loeblich & Loeblich, 1966 [Treated 
by Lentin & Williams, 1981, as a subspe- 
cies] 

Spiniferites ramosus var. reticulatus (Davey & 
Williams, 1966a) Davey & Verdier, 1971 
[Treated by Lentin & Williams, 198 1, as 
a subspecies] 



Litosphaeridium arundum (Eisenack & Cook- 
son, i960) Davey, 1979 



Florentinia clavigera (Deflandre, 1937) Davey 
& Verdier, 1973; emend. Davey & Verdier, 
1976 

Compositosphaeridium costatum (Davey & 
Williams, 1966b) Dodekova, 1974 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 



39 



HYSTRICHOSPHAERIDIUM continued 
deanei Davey & Williams, 1966b 

Pgliwicense Macko 1957 

hillii (Merrill, 1895) Sarjeant, 1964 

irregulare (Merrill, 1895) Sarjeant, 1964 



patulum Davey & Williams, 1966b 
polyplasium Maier, 1959* 
radiculatum Davey & Williams, 1966b 

readei Davey & Williams, 1966b 

salpingophorum Deflandre. 1935; emend. 

Davey & Williams, 1966b 
sheppeyense Davey & Williams, 1966b 



sitnplicispinum Davey & Williams, 1966b 

stellatum Maier, 1959* 
tridactylites Valensi, 1955 

tubiferum (Ehrenberg, 1838) Deflandre, 
1937; emend. Davey & Williams, 1966b 

tubiferum var. brevispinum Davey & 
Williams, 1966b 

HYSTRICHOSPHAEROPSIS Deflandre, 
1935; emend. Sarjeant, 1966b; emend. 
Gocht, 1976 
borussica Eisenack, 1954 

ovum Deflandre, 1935 

wetzeli (Deflandre, 1937) Sarjeant, 1966b 



Florentinia deanei (Davey & Williams, 1966b) 
Davey & Verdier, 1973 



Hystrichosphaeridium speciale (Merrill, 1895) 
Lentin & Williams, 1981. [At the time of 
the transfer, this name was a jr horn, of 
Hystrichosphaeridium irregulare Pocock, 
1962; however, the latter species was 
transferred to Oligosphaeridium by Davey 
& Williams in Davey et al., 1969. Never- 
theless, Lentin & Williams (1981) con- 
sidered that the name H. irregulare (Mer- 
rill) must still be considered a. jr horn, and 
accordingly proposed the above substi- 
tute name. Though Art. 55 of the Inter- 
national Code of Botanical Nomenclature 
would require reinstatement of the name 
irregulare (Merrill) if that species be trans- 
ferred to any genus other than Oligo- 
sphaeridium, their procedure appears 
correct.] 



Florentinia radiculata (Davey & Williams, 
1966b) Davey & Verdier, 1976 

Kleithriasphaeridium readei (Davey & 
Williams, 1966b) Davey & Verdier, 1976 



Shown by Stover & Evitt, 1978, to be &jr syn. 
of Hystrichosphaeridium pseudorecurva- 
tum Morgenroth, now M elitasphaeridium 
pseudorecurvatum (Morgenroth, 1966) 
Bujak in Bujak et al., 1980. 

Kleithriasphaeridium simplicispinum (Davey 
& Williams, 1966b) Davey, 1974 

Florentinia tridactylites (Valensi, 1955) Dux- 
bury, 1980 



[Treated as a subspecies by Lentin & 
Williams, 1973.] 



Rottnestia borussica (Eisenack, 1954) Cookson 
& Eisenack, 1961 

Spiniferites wetzelii (Deflandre, 1937) Sarjeant, 
1970 



4 o 



SECOND APPENDIX TO 



LEPTODINIUM Klement, i960; emend. 
Sarjeant, 1982a 
aceras (Eisenack, 1958) Sarjeant, 1969 

aculeatum Wall, 1967 

alectrolophum Sarjeant, 1966b 

amabilis (Deflandre, 1939) Sarjeant, 1969 

arcuatum Klement, i960** 

clathratum (Cookson & Eisenack, 1960b) 

Sarjeant, 1969 
crassinervum (Deflandre, 1939) Sarjeant, 

1969 
Pdelicatum (Davey, 1969a) Sarjeant, 1969 
eumorphum (Cookson & Eisenack, 1960b) 

Sarjeant, 1969 
freakei (Sarjeant, 1963) Sarjeant, 1969 
maculatum Cookson & Eisenack, 1961 

margaritiferum (Cookson & Eisenack, 1961) 

Sarjeant, 1969 
membranigerum Gerlach, 1961 

millioudi (Sarjeant, 1963) Sarjeant, 1969 
mirabile Klement, i960** 
mosaicum (Downie, 1957) Sarjeant, 1969 
paradoxum Wall, 1967 

sphaericum Wall, 1967 

strialatum Wall, 1967 

striatum (Clarke & Verdier, 1967) Sarjeant, 

1969 
subtile Klement, i960 
Ptenuicornutum Cookson & Eisenack, 1962 

LITOSPHAERIDIUM Davey & Williams, 
1966b; emend. Davey & Verdier, 1973 

Pcrassipes (Reade, 1839) Davey & Williams, 
1969 

Pflosculus (Deflandre, 1937) Davey & 
Williams, 1969 

Pinversibuccinum Davey & Williams, 1966b 

siphoniphorum (Cookson & Eisenack, 1958) 

Davey & Williams, 1966b 
Ptruncigerum (Deflandre, 1937) Davey & 

Williams, 1969 

MEIOUROGONYA ULAX Sarjeant, 1966b 
bulloidea (Cookson & Eisenack, 1960b) Sar- 
jeant, 1969 
caytonensis (Sarjeant, 1959) Sarjeant, 1969 



AcanthaulaxP aceras (Eisenack, 1958) Stover 

&Evitt, 1978 
Impagidinium aculeatum (Wall, 1967) Lentin 

& Williams, 1981 
Impagidinium alectrolophum (Sarjeant, 1966b) 

Stover & Evitt, 1978 

Leptodinium amabile (Deflandre, 1939) Sar- 
jeant, 1969 



Impagidinium maculatum (Cookson & Eisen- 
ack, 1961) Stover & Evitt, 1978 

Impagidinium margaritiferum (Cookson & 
Eisenack, 1961) Stover & Evitt, 1978 

Impagidinium? membranigerum (Gerlach, 
1961) Stover & Evitt, 1978 



Impagidinium paradoxum (Wall, 1967) Stover 

& Evitt, 1978 
Impagidinium sphaericum (Wall, 1967) Lentin 

& Williams, 1981 
Impagidinium strialatum (Wall, 1967) Stover 

& Evitt, 1978 
Impagidinium striatum (Clarke & Verdier, 

1967) Stover & Evitt, 1978 



Florentinia? flosculus (Deflandre, 1937) Lentin 

& Williams, 1981 
Paucisphaeridium inversibuccinum (Davey & 

Williams, 1966b) Bujak et al., 1980 



Cordosphaeridium truncigerum (Deflandre, 
1937) De Coninck, 1975 






DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 



4i 



MEIOUROGONYAULAX continued 
Pcristulata (Sarjeant, 1959) Sarjeant, 1969 
decapitata (W. Wetzel, 1967) Sarjeant, 1969; 

emend. Sarjeant, 1980 
superornata (W. Wetzel, 1967) Sarjeant, 

1969; emend. Sarjeant, 1980 
valensii Sarjeant, 1966b 

MEMBRANILARNACIA Eisenack, 1963a; 

emend. Williams & Downie, 1966c 
leptoderma (Cookson & Eisenack, 1958) 

Eisenack, 1963a 
reticulata Williams & Downie, 1966c 



MICRODINIUM Cookson & Eisenack, 
1960a; emend. Sarjeant, 1966b; emend. 
Stover & Evitt, 1978 

ornatum Cookson & Eisenack, 1960a 

setosum Sarjeant, 1966b 

MUDERONGIA Cookson & Eisenack, 1958 
mcwhaei Cookson & Eisenack, 1958 
staurota Sarjeant, 1966c 

NEMATOSPHAEROPSIS Deflandre & 
Cookson, 1955; emend. Williams & 
Downie, 1966c 
balcombiana Deflandre & Cookson, 1955 

NETRELYTRON Sarjeant, 1961; emend. 
Sarjeant, 1966c 



jurassicum (Alberti, 1961) Sarjeant, 1966c 



stegastum Sarjeant, 1961; emend. Sarjeant, 
1966c 



trinetron Sarjeant, 1966c 



ODONTOCHITINA Deflandre, 1935; emend. 
Davey, 1970 
operculata (O. Wetzel, 1933) Deflandre & 
Cookson, 1955 

OLIGOSPHAERIDIUM Davey & Williams, 
1966b 
albertense (Pocock, 1962) Davey & Williams, 

1969 
anthophorum (Cookson & Eisenack, 1958) 
Davey, 1969a 



[Considered by De Coninck, 1969 and Gocht, 
1969 to be a jr syn. of M embranilarnacia 
ursulae (Morgenroth, 1966) De Coninck, 
1969 now Eatonicysta ursulae (Morgen- 
roth, 1966) Stover & Evitt, 1978.] 



[Wiggins, 1975, considered this a jr syn. of 

Kalyptea Cookson & Eisenack, 1960b. 

This judgement was rejected by Sarjeant, 

1976] 
Kalyptea jurassica Alberti, 1961. [Since this 

species was not validly transferred in 1966 

(see Lentin & Williams, 1981) it must be 

retained in Kalyptea.'] 
[The transfer of this species to Kalyptea by 

Wiggins, 1975, was rejected by Sarjeant, 

1976.] 
[The transfer of this species to Kalyptea by 

Wiggins, 1975, was rejected by Sarjeant, 

1976] 



42 



SECOND APPENDIX TO 



OLIGOSPHAERIDIUM continued 

Pasterigerum (Gocht, 1959) Davey & 

Williams, 1969 
Pcoelenteratum (Tasch, 1964) Davey & 
Williams, 1969 

complex (White, 1842) Davey & Williams, 

1969 
dictyophorum (Cookson & Eisenack, 1958) 

Davey & Williams, 1969 
Pdispare (Tasch, 1964) Davey & Williams, 

1969 

Pirregulare (Pocock, 1962) Davey & 

Williams, 1969 
macrotubulum (Neale & Sarjeant, 1962) 

Davey & Williams, 1966b 



Pparadoxum (Brosius, 1963) Davey & 

Williams, 1969 
Pperforatum (Gocht, 1959) Davey & 

Williams, 1969 
prolixospinosum Davey & Williams, 1966b 
pulcherrimum (Deflandre & Cookson, 1955) 

Davey & Williams, 1966b 
reniforme (Tasch, 1964) Davey, 1969a 



reticulatum Davey & Williams, 1966b 
vasi/ormum (Neale & Sarjeant, 1962) Davey 
& Williams, 1966b 



Shown by Stover & Evitt, 1978, to be a.jr syn. 
of Oligosphaeridium irregular e (Pocock, 
1962) Davey & Williams, 1969 



Shown by Stover & Evitt, 1978, to be a.jr syn. 
of Oligosphaeridium irregular e (Pocock, 
1962) Davey & Williams, 1969 



[Considered by Mclntyre & Brideaux, 1980, 
to be a.jr syn. of Oligosphaeridium vasi/or- 
mum (Neale & Sarjeant, 1962) Davey & 
Williams, 1966b; the latter species is in 
turn considered a jr syn. of Oligosphaeri- 
dium asterigerum (Gocht, 1959) Davey & 
Williams, 1969, by Duxbury, 1977.] 
Bipolaribucina paradoxa (Brosius, 1963) 
"Jiabo" 1978 



Shown by Stover & Evitt, 1978, to be a jr 
syn. of Oligosphaeridium irregular e (Po- 
cock, 1962) Davey & Williams, 1969. 

[Considered by Duxbury, 1977, to be a 
jr syn. of Oligosphaeridum asterigerum 
(Gocht, 1959) Davey & Williams, 1969. 
This judgment is not accepted by us.] 



PARANETRELYTRON Sarjeant, 1966c 



strongylum Sarjeant, 1966c 



[Considered by Wiggins, 1975, to be ajr syn. 
of Pareodinia Deflandre, 1947. This judge- 
ment was rejected by Sarjeant, 1976.] 

[Considered by Wiggins, 1975, to be ajr syn. 
of Pareodinia ceratophora Deflandre, 1947. 
This judgement was rejected by Sarjeant, 
1976.] 



PAREODINIA Deflandre, 1947; emend. Wig- 
gins, 1975; emend. Stover & Evitt, 1978 
ceratophora Deflandre, 1947; emend. Gocht, 
1970 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 



43 



PERISSEIASPHAERIDIUM Davey & 
Williams, 1966b 
eisenackii (Eisenack, 1958) Davey & 
Williams, 1969 

pannosum Davey & Williams, 1966b 
PLURIARVALIUM Sarjeant, 1962a 



osmingtonense Sarjeant, 1962b 

POLYSPHAERIDIUM Davey & Williams, 
1966b; emend. Bujak et al., 1980 
Paspetum (Maier, 1959) Davey & Williams, 

1969* 
Pbelgicum Sarjeant, 1969 

Pdeflandrei (Valensi, 1947) Davey & 

Williams, 1969 
Pfabium (Tasch, 1964) Davey & Williams, 

1969 

Pfluctuans (Pastiels, 1948) Davey & Williams 
1966b 

Pfollium (Tasch, 1964) Davey & Williams, 
1969 

Pfucosum (Valensi, 1955) Davey & Williams, 

1969 
laminaspinosum Davey & Williams, 1966b 

Pmajor (Lejeune-Carpentier, 1940) Davey & 

Williams, 1969 
Pmarsupium (Tasch, 1964) Davey & 

Williams, 1969 

pastielsi Davey & Williams, 1966b 

Ppaulinae (Valensi, 1953) Davey & Williams, 

1969 
Pperovatum (Tasch, 1964) Davey & Williams, 

1969 

Ppolypes (Cookson & Eisenack, 1962) Davey 

& Williams, 1966b 
pumilum Davey & Williams, 1966b 

Prhabdophorum (Valensi, 1955) Davey & 

Williams, 1969 
Psimplex (White, 1842) Davey & Williams, 

1969 



[Considered by Davey & Verdier, 1974 and 
Duxbury, 1980, to be &jr syn. of Hystrich- 
osphaerina schindewolfii Alberti, 1961.] 

[Considered by Wiggins, 1975 and Dorhofer & 
Davies, 1980 to be a.jt syn. of Pareodinia 
Deflandre, 1947. We concur with Stover 
& Evitt, 1978, who neither accept this 
judgement nor the resultant transfer of 
the type species.] 



Dapsilidinium asperum (Maier, 1959) Bujak et 
al., 1980* 

Dapsilidinium? belgicum (Sarjeant, 1969) Len- 
tin & Williams, 1981 

DapsilidiniumP deflandrei (Valensi, 1947) Len- 
tin & Williams, 1981 

Shown by Stover & Evitt, 1978, to be a syn- 
onym of Kiokansiutn unituberculatum 
(Tasch, 1964) Stover & Evitt, 1978 

The taxonomic transfer of this species was not 
validly made; therefore it must remain in 
H ystrichosphaeridium . 

Shown by Stover & Evitt, 1978, to be a syn- 
onym of Kiokansiutn unituberculatum 
(Tasch, 1964) Stover & Evitt, 1978. 

Dapsilidinium? fucosum (Valensi, 1955) Lentin 
& Williams, 1981 

Dapsilidinium laminaspinosum (Davey & 
Williams, 1966b) Lentin & Williams, 1981 

Dapsilidinium? major (Lejeune-Carpentier, 
1940) Lentin & Williams, 1981 

Shown by Stover & Evitt, 1978, to be a syn- 
onym of Kiokansiutn unituberculatum 
(Tasch, 1964) Stover & Evitt, 1978. 

Dapsilidinium pastielsii (Davey & Williams, 
1966b) Bujak et al., 1980 

Dapsilidinium? paulinae (Valensi, 1953) Len- 
tin & Williams, 1981 

Shown by Stover & Evitt, 1978, to be a syn- 
onym of Kiokansiutn unituberculatum 
(Tasch, 1964) Stover & Evitt, 1978. 

Bacchidinium polypes (Cookson & Eisenack, 
1962) Davey, 1979 

Dapsilidinium? pumilum (Davey & Williams, 
1966b) Lentin & Williams, 1981 

Dapsilidinium? thabdophotum (Valensi, 1955) 
Lentin & Williams, 1981 

Dapsilidinium simplex (White, 1842) Bujak et 
al., 1980 



44 



SECOND APPENDIX TO 



POLYSPHAERIDIUM continued 

subtile Davey & Williams, 1966b; emend. 

Bujak et al., 1980 
?tribrachiosum (Tasch, 1964) Davey & 
Williams, 1969 

zoharyi (Rossignol, 1962) Bujak et al., 1980 

PROLIXOSPHAERIDIUM Davey et al., 
1966 
deirense Davey et al., 1966 



granulosum (Deflandre, 1937) Davey et al., 

1966 
mixtispinosum (Klement, i960) Davey et al., 

1969** 
parvispinutn (Deflandre, 1937) Davey et al., 

1969 
? xanthiopyxid.es (O. Wetzel, 1933) Davey et 

al., 1969 

PSALIGONYAULAX Sarieant, 1966b 
apatela (Cookson & Eisenack, 1960b) Sar- 
jeant, 1969 



deflandrei Sarjeant, 1966b 
simplicia (Cookson & Eisenack, 1961) Sar- 
jeant, 1969 

RAPHIDODINIUM Deflandre, 1936 
fucatum Deflandre, 1936 

RHAETOGONYAULAX Sarjeant, 1966b; 
emend. Harland et al., 1975; emend. 
Fisher & van Helden, 1979 
chaloneri (Sargeant, 1963) Sarjeant 1969 

rhaetica (Sarjeant, 1963) Loeblich & Loe- 
blich, 1968; emend. Harland et al., 1975 

RHYNCHODINIOPSIS Deflandre, 1935; 
emend. Below, 1981; emend. Sarjeant, 
1982a 
aptiana Deflandre, 1935; emend. Sarjeant, 
1982a 

SIRMIODINIUM Alberti, 1961; emend. 
Warren, 1973 
gross* Alberti, 1961; emend. Warren, 1973 

SURCULOSPHAERIDIUM Davey et al., 
1966 
cribrotubiferum (Sarjeant, i960) Davey etal., 
1966 



Shown by Stover & Evitt, 1978, to be a syn- 
onym of Kiokansium unituberculatum 
(Tasch, 1964) Stover & Evitt, 1978. 



[Considered by Davey & Verdier, 1974, to be 
a jr syn. of Prolixosphaeridium parvi- 
spinutn (Deflandre, 1937) Davey et al., 
1969.] 



Tanyosphaeridium xanthiopyxides (O. Wetzel, 
1933) Stover & Evitt, 1978 

Tubotuberella apatela (Cookson & Eisenack, 
1960b) Ioannides et al., 1976; emend. Sar- 
jeant, 1982a. [The previous placement of 
this species as the type for the new genus 
Glabridinium by Brideaux, 1977, was re- 
jected by Sarjeant, 1982a] 

Impagidinium? simplicium (Cookson & Eisen- 
ack, 1961) Stover & Evitt, 1978 



Rhaetogonyaulax rhaetica subsp. chaloneri (Sar- 
jeant, 1963) Lentin & Williams, 1977 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 



45 



SURCULOSPHAERIDIUM continued 
longifurcatum (Firtion, 1952) Davey et al., 

1966 
vestitum (Deflandre, 1938) Davey et al., 1966 

SYSTEMATOPHORA Klement, i960 
areolata Klement, i960 
placacantha (Deflandre & Cookson, 1955) 

Davey et al., 1966; emend. May, 1980 
schindewolfi (Alberti, 1961) Downie & Sar- 
jeant, 1965 



TANYOSPHAERIDIUM Davey & 

Williams, 1966b 
ellipticum (Cookson, 1965) Davey & 

Williams, 1969 
isocalamum (Deflandre & Cookson, 1955) 

Davey & Williams, 1969 
regulare Davey & Williams, 1966b 
variecalamum Davey & Williams, 1966b 

THALASSIPHORA Eisenack & Gocht, i960; 
emend. Gocht, 1968 
delicata Williams & Downie, 1966c; emend. 

Eaton, 1976 
pelagica (Eisenack, 1954) Eisenach & Gocht, 
i960 

WANAEA Cookson & Eisenack, 1958; emend. 
Fensome, 1981 
spectabilis (Deflandre & Cookson, 1955) 
Cookson & Eisenack, 1958 

WETZELIELLA Eisenack, 1938; emend. 
Williams & Downie, 1966b; emend. Lentin 
& Williams, 1976 

articulata Eisenack, 1938 
articulata var. conopia Williams & Downie, 
1966b 

clathrata Eisenack, 1938 

coleothrypta Williams & Downie, 1966b 

condylos Williams & Downie, 1966b 

homomorpha Deflandre & Cookson, 1955 



homomorpha var. quinquelata Williams & 

Downie, 1966b 
ovalis Eisenack, 1954 



Hystrichosphaerina schindewolfii Alberti, 1961. 
[The genus Hystrichosphaerina, rejected 
as a.jr syn. of Systematophora by Neale & 
Sarjeant (1962), was reinstated under an 
emended definition by Stover & Evitt, 
1978.] 



Distatodinium ellipticum (Cookson, 1965) 
Eaton, 1976 



[Since the other constituent subgenera have 
either been raised to generic status or 
abandoned, the type subgenus Wetzeliella 
has now fallen into disuse.] 

Kisselovia tenuivirgula subsp. conopia 

(Williams & Downie, 1966b) Costa & 

Downie, 1979 
Kisselovia? clathrata (Eisenack, 1938) Lentin 

& Williams, 1976 
Kisselovia coleothrypta (Williams & Downie, 

1966b) Lentin & Williams, 1976 
Dracodinium condylos (Williams & Downie, 

1966b) Costa & Downie, 1979 
Apectodinium homomorphum (Deflandre & 

Cookson, 1955) Lentin & Williams, 1977; 

emend. Harland, 1979 
Apectodinium quinquelatum (Williams & 

Downie, 1966b) Costa & Downie, 1979 



46 



SECOND APPENDIX TO 



WETZEL1ELLA continued 

reticulata Williams & Downie, 1966b 

similis Eisenack, 1954 

solida (Gocht, 1955) Williams & Downie, 

1966b 
symmetrica Weiler, 1956 
symmetrica var. lobisca Williams & Downie, 

1966b 

tenuivirgula Williams & Downie, 1966b 

tenuivirgula var. crassiramosa Williams & 

Downie, 1966b 
varielongituda Williams & Downie, 1966b 

WETZELIELLA (RHOMBODINIUM) 
Gocht, 1955 
draco Gocht, 1955 
glabra Cookson, 1956 

XIPHORIDIUM Sarjeant, 1966b 

alatum (Cookson & Eisenack, 1962) Sar- 
jeant, 1966b 



Kisselovia reticulata (Williams & Downie, 
1966b) Lentin & Williams, 1976 

Dracodinium simile (Eisenack, 1954) Costa & 
Downie, 1979 

Dracodinium solidum Gocht, 1955 



Wetzeliella meckelfeldensis subsp. lobisca 

(Williams & Downie, 1966b) Costa & 

Downie, 1979 
Kisselovia tenuivirgula (Williams & Downie, 

1966b) Lentin & Williams, 1976 
Kisselovia crassiramosa (Williams & Downie, 

1966b) Damassa, 1979 
Dracodinium varielongitudum (Williams & 

Downie, 1966b) Costa & Downie, 1979 

Raised to generic status by Gocht, 1955; 

emend. Bujak, 1979. 
Rhombodinium draco Gocht, 1955 
Rhombodinium glabra (Cookson, 1956) Voz- 

zhennikova, 1967 



REFERENCES 



Alberti, G. 1959. Zur Kenntnis der Gattung Deflandrea Eisenack (Dinoflag.) in der Kreide und im 

Alttertiar Nord- und Mitteldeutschlands. Mitt. geol. Stlnst. Hamb. 28: 93-105, pis 8, 9. 
1961. Zur Kenntnis mesozoischer und alttertiarer Dinoflagellaten und Hystrichosphaerideen 

von Nord- und Mitteldeutschland sowie einigen anderen Europaischen Gebieten. Palaeontogra- 

phica, Stuttgart, (A) 116: 1-48, pis 1-12. 
Baksi, S. K. 1962. Palynological investigation of Simsang River Tertiaries, south Shillong Front, 

Assam. Bull. geol. min. metall. Soc. India, Calcutta, 26: 1-22, pis 1-5. 
Below, R. 1981. Dinoflagellaten-zysten aus dem oberen Hauterive bis unteren Cenoman Siid- 

West-Marokkos. Palaeontographica, Stuttgart, (B) 176: 1-145, pis 1-15. 
1982. Rigaudella, ein neues Genus von Dinoflagellaten-zysten. Neues Jb. Geol. Paldont. Mh., 

Stuttgart, 1983 (3): 137-150, text-figs 1-34. 
Brideaux, W. W. 1977. Taxonomy of Upper Jurassic-Lower Cretaceous microplankton from the 

Richardson Mountains, District of Mackenzie, Canada. Bull. geol. Surv. Can., Ottawa, 281: 1- 

89, pis 1-16. 
— & Fisher, M. J. 1976. Upper Jurassic-Lower Cretaceous dinoflagellate assemblages from 

Arctic Canada. Bull. geol. Surv. Can., Ottawa, 259: 1-53, pis 1-17. 
Brosius, M. 1963. Plankton aus dem nordhessischen KasselerMeeressand (Oberoligozan).Z. dt. Geol. 

Ges., Berlin, 1 14: 32-56, pis 1-8. 
Bujak, J. P. 1979. Proposed phylogeny of the dinoflagellates Rhombodinium and Gochtodinium. 

Micropaleontology , New York, 25: 308-324, pis 1-3. 
, Downie, C, Eaton, G. L. & Williams, G. L. 1980. Taxonomy of some Eocene dinoflagellate 

cyst species from southern England. In Bujak, J. P., Downie, C, Eaton, G. L. & Williams, 

G. L., Dinoflagellate cysts and acritarchs from the Eocene of southern England. Spec. Pap. 

Palaeont., London. 25: 26-36. 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 47 

Churchill, D. M. & Sarjeant, W. A. S. 1963. Freshwater microplankton from Flandrian (Holo- 

cene) peats of south-western Australia. Grana palynol., Stockholm, 3: 29-53, pis 1-2. 
Clarke, R. F. A. & Verdier, J. -P. 1967. An investigation of microplankton assemblages from the 

Chalk of the Isle of Wight, England. Verh. K. ned. Akad. Wet., Amsterdam, 24 (3): 1-96, pis 

1-17. 
Cookson, I. C. 1956. Additional microplankton from Australian Late Mesozoic and Tertiary 

sediments. Aust. J . mar. Freshwat. Res., Melbourne, 7: 183-191, pis 1-2. 

1965. Cretaceous and Tertiary microplankton from southeastern Australia. Proc. R. Soc. 

Victoria, Melbourne, 78: 85-93, pis 9-11. 

& Eisenack, A. 1958. Microplankton from Australian and New Guinea Upper Mesozoic 

sediments. Proc. R. Soc. Victoria, Melbourne, 70: 19-79, P' s 1-12. 

1960a. Microplankton from Australian Cretaceous sediments. M icropaleontology , New 
York, 6: 1-18, pis 1-3. 

1960b. Upper Mesozoic microplankton from Australia and New Guinea. Palaeontology, 

London, 2: 243-261, pis 37-39. 

1961. Tertiary microplankton from the Rottnest Island Bore, Western Australia. // R. 



Soc. West. Aust., Perth, 44: 39-47, pis 1-2. 

1962. Additional microplankton from Australian Cretaceous sediments. Micropaleon- 
tology, New York, 8: 485-507, pis 1-7. 

1968. Microplankton from two samples from Gingin Brook No. 4 Borehole, Western 
Australia. Jl R. Soc. West. Aust., Perth, 51: 110-122. 

1974. Mikroplankton aus Australischen Mesozoischen und Tertiaren Sedimenten. Palaeon- 



tographica, Stuttgart, (B) 148: 44-93, pis 20-29. 
Corradini, D. 1973. Non-calcareous microplankton from the Upper Cretaceous of the Northern 

Apennines. Boll. Soc. paleont. ital., Modena, II: 119- 197, pis 19-39. 
Costa, L. I. & Downie, C. 1979. The Wetzeliellaceae; Palaeogene dinoflagellates. Proc. IV int. 

Palynol. Con/., Lucknow, (1976-77) 2: 34-46. 
Damassa, S. P. 1979. Eocene dinoflagellates from the Coastal Belt of the Franciscan Complex, 

northern California. /. Paleont., Tulsa, 53: 815-840, pis 1-8. 
Davey, R. J. 1968. Gonyaulacysta parorthoceras, a new species of dinoflagellate cyst. Palaeont. a/r., 

Johannesburg, 11: 1. 
1969a. Non-calcareous microplankton from the Cenomanian of England, northern France and 

North America, Part I. Bull. Br. Mus. nat. Hist., London, (Geol.) 17: 103-180, pis 1-11. 
1969b. Some dinoflagellate cysts from the Upper Cretaceous of northern Natal, South Africa. 

Palaeont. afr., Johannesburg, 12: 1-23, pis 1-4. 
1969c. The evolution of certain Upper Cretaceous hystrichospheres from South Africa. 

Palaeont. afr., Johannesburg, 12: 25-51, pis 1-4. 
1970. Non-calcareous microplankton from the Cenomanian of England, northern France and 

North America, Part II. Bull. Br. Mus. nat. Hist., London, (Geol.) 18: 333-397, pis 1-10. 
T 974 Dinoflagellate cysts from the Barremian of the Speeton Clay, England. In: Symposium 

on Stratigraphic Palynology. Spec. Pubis Birbal Sahni Inst. Palaeobot., Lucknow, 3: 41-75, pis 

1-9. 
1979- Marine Apto-Albian palynomorphs from Holes 400A and 402A, IPOD Leg 48, northern 

Bay of Biscay. Initial Rep. deep Sea Drilling Proj., Washington, 48: 547-577, pis 1-8. 
, Downie, C, Sarjeant, W. A. S. & Williams, G. L. 1966. Fossil dinoflagellate cysts attributed 

to Baltisphaeridium. In Davey, R. J., Downie, C, Sarjeant, W. A. S. & Williams, G. L., Studies 

on Mesozoic and Cainozoic dinoflagellate cysts. Bull. Br. Mus. nat. Hist., London, (Geol. Suppl.) 

3:157-175. 
1969. Generic reallocations. In Davey, R. J., Downie, C, Sarjeant, W. A. S. 

& Williams, G. L. Appendix to "Studies on Mesozoic and Cainozoic dinoflagellate cysts". Bull. 

Br. Mus. nat. Hist., London, (Geol. Suppl.) 3 Appendix: 15-17. 
& Verdier, J. -P. 1971. An investigation of microplankton assemblages from the Albian of the 

Paris Basin. Verh. K. ned. Akad. Wet., Amsterdam, 26: 1-58, pis 1-7. 

1973. An investigation of microplankton assemblages from latest Albian (Vraconian) 

sediments. Revta esp. Micropaleont., Madrid, 5: 173-212, pis 1-5. 



48 SECOND APPENDIX TO 

Davey, R. J. & Verdier, J. -P. 1974. Dinoflagellate cysts from the Aptian type sections at Gargas 
and La Bedoule, France. Palaeontology , London, 17: 623-653, pis 91-93. 

1976. A review of certain non-tabulate Cretaceous hystrichospherid dinocysts. Rev. 
Palaeobot. Palynol., Amsterdam, 22: 307-335, pis 1-4. 
- & Williams, G. L. 1966a. The genera Hystrichosphaera and Achomosphaera. In Davey, R. J., 
Downie, C, Sarjeant, W. A. S. & Williams, G. L. Studies on Mesozoic and Cainozoic dinofla- 
gellate cysts. Bull. Br. Mus. nat. Hist., London, (Geol. Suppl.) 3: 28-52. 

1966b. The genus Hystrichosphaeridium and its allies. In Davey, R. J., Downie, C, 
Sarjeant, W. A. S. & Williams, G. L. Studies on Mesozoic and Cainozoic dinoflagellate cysts. 
Bull. Br. Mus. nat. Hist., London, (Geol. Suppl.) 3: 53-106. 

1969 Generic reallocations. In Davey, R. J., Downie, C, Sarjeant, W. A. S. & Williams, 

G. L. Appendix to "Studies on Mesozoic and Cainozoic dinoflagellate cysts". Bull. Br. Mus. 
nat. Hist., London, (Geol. Suppl.) 3 Appendix: 4-7. 

De Coninck, J. 1969. Dinophyceae et Acritarcha de l'Ypresien du Sondage de Kallo. Mem. Inst. r. 
Sci. nat. Belg., Brussels, 161: 1-67, pis 1-17. 

1975- Microfossiles a paroi organique de l'Ypresien du Bassin beige. Prof. Pap. belg. geol. Dienst, 

Brussels, 12: 1-151, pis 1-22. 

Deflandre, G. 1935. Considerations biologiques sur les microorganismes d'origine planctonique 
conserves dans les silex de la craie. Bull. biol. Fr. Belg., Paris, 69: 213-244, pis 5-9. 

1936. Microfossiles des silex cretaces. I. Generalites. Flagelles. Annls Paleont., Paris, 25: 151 -191 

(3-43). pis 11-20(1-10). 

1937- Microfossiles des silex cretaces. II. Flagelles incertae sedis. Hystrichosphaerides. Sarco- 

dines. Organismes divers. Annls Paleont., Paris, 26: 51-103 (3-55), pis 8-15 (11-18). 

1938. Microplancton des mers jurassiques conserve dans les marnes de Villers-sur-Mer (Calva- 
dos). Trav. Stn zool. Wimereux, Paris, 13: 147-200, pis 5-11. 

1939- Sur les dinoflagelles des schistes bitumineux d'Orbagnoux (Jura). Bull. Soc. fr. Microsc, 

Paris, 8: 141-145, pi. 6. 

1946. Hystrichosphaerides II. Gymnodiniales et Dinoflagelles incertae sedis. Flagelles incertae 

sedis. Fichier micropaleont. (6), Arch. orig. Serv. Docum. C. N. R. S., 235: cards 860-1019. 

1947 Sur quelques microorganismes planctoniques des silex Jurassiques. Bull. Inst, oceanogr. 

Monaco, 921: 1-10, figs 1-5. 

1954- Systematique des Hystrichosphaerides: sur l'acception du genre Cymatiosphaera O. 

Wetzel. C . r. somm. Seanc. Soc. geol. France, Paris, 1954 (12): 257-258. 

1964. Remarques sur la classification des Dinoflagelles fossiles, a propos d' Evittodinium , 

nouveau genre cretace de la famille des Deflandreaceae. C.r. hebd. Seanc. Acad. Sci., Paris, 258: 
5027-5030. 

& Cookson, I.C. 1955. Fossil microplankton from Australian Late Mesozoic and Tertiary 

sediments. Aust. J. mar. Freshwat. Res., Melbourne, 6: 242-313, pis 1-9. 
& Courteville, H. 1939. Note preliminaire sur les microfossiles des silex cretaces du Cambresis. 

Bull. Soc. fr. Microsc, Paris, 8: 95-106, pis 1-3. 
& Deflandre-Rigaud, M. 1958. Hysterichosphaerides IV et genres incertae sedis. Especes du 

Secondaire et du Tertiaire (Supplement 1). Fichier micropaleont. (10), Arch. orig. Serv. Docum. 

C. N. R. S., 366: cards 1293-1750. 
Deunff, J. 1961. Un microplankton a Hystrichospheres dans leTremadocdu Sahara. Revue Micro- 
paleont., Paris, 4: 37-52, pis 1-3. 
De Wit, R. 1943. Hystrichosphaeridae uit Limburgsche Vuursteen. Verh. geol.-mijn. Genoot. Ned., 

s'Gravenhage, 13: 363-392. 
Dodekova, L. 1967. Les dinoflagelles et acritarches de 1'Oxfordien-Kimeridgien de la Bulgarie du 

Nord-Est. God. sof. Univ., 60 (1, Geol.): 9-30, pis 1-3. 
1974. Compositosphaeridium gen. n. - a new genus dinoflagellate cyst. Izv. geol. Inst., Sofia, 

(ser. pal.) 23: 25-30, pis 1-2. 
1975- New Upper Bathonian dinoflagellate cysts from northeastern Bulgaria. Paleont. Stratig. 

Litolog., Sofia, 2: 17-34, pis 1-6. 
Dorhofer, G. & Davies, E. H. 1980. Evolution of archeopyle and tabulation in Rhaetogonyaulacinean 

dinoflagellate cysts. 91 pp., 40 figs. Toronto (Life Sci. misc. Pubis R. Ont. Mus.) 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 49 

Downie, C. 1957. Microplankton from the Kimeridge Clay. Q. Jl geol. Soc. Lond., 112: 413-434, 

pi. 20. 
& Sarjeant, W. A. S. 1963. On the interpretation and status of some hystrichosphere genera. 

Palaeontology, London, 6: 83-96. 
1965. Bibliography and index of fossil dinoflagellates and acritarchs. Mem. geol. Soc. Am., 

Washington, 94: 1-180. 
Drugg, W. S. 1964. Glyphanodinium, a new dinoflagellate genus from the Paleocene of California. 

Proc. biol. Soc. Wash., 77: 237-240, 6 figs. 

1967. Palynology of the Upper Moreno Formation (Late Cretaceous-Paleocene) Escarpado 

Canyon, California. Palaeontographica, Stuttgart, (B) 120: 1-71, pis 1-9. 

Dupin, F. 1968. Deux nouvelles especes de Dinoflagelles du Jurassique d'Aquitaine. Cah. Micro- 

paleont., Paris, (ser. I) 8: 1-5, pi. 1. 
Duxbury, S. 1977. A palynostratigraphy of the Berriasian to Barremian of the Speeton Clay of 

Speeton, England. Palaeontographica, Stuttgart, (B) 160: 17-67, pis 1-15. 
1980. Barremian phytoplankton from Speeton, east Yorkshire. Palaeontographica , Stuttgart, 

(B) 173: 107-146, pis 1-13. 
Eaton, G. L. 1971. A morphogenetic series of dinoflagellate cysts from the Bracklesham Beds of the 

Isle of Wight, Hampshire, England. In Farinacci, A. (ed.), Proc. II Planktonic Conference, 

Roma igyo, 1: 355-379, pis 1-4. Rome. 
1976. Dinoflagellate cysts from the Bracklesham Beds (Eocene) of the Isle of Wight, Southern 

England. Bull. Br. Mus. nat. Hist., London, (Geol.) 26: 227-332, pis 1-21. 
Ehrenberg, C. G. 1838. Uberdas Massenverhaltniss der jetzt lebenden Kiesel-Infusorien und fiber ein 

neues Infusorien-Conglomerat als Polirschiefer von Jastraba in Ungarn. Phys. Math. Abh. K. 

Akad. Wiss. Berlin., 1836: 109-135, pis 1-2. 
1843. Uber einige Jura-Infusorien-Arten des Corallrags bei Krakau. Ber. Akad. Wiss. Berlin, 

1843:61-63. 
Eisenack, A. 1935. Mikrofossilien aus Doggergeschieben Ostpreussens. Z. Geschiebeforsch. , Berlin, 

11: 167-184, pis 4-5. 

1938. Hystrichosphaerideen und verwandte Formen in baltischen Silur. Z. Geschiebeforsch., 

Berlin, 14: 1-30, pis 1-4, text-figs 1-7. 

1954- Mikrofossilien aus Phosphoriten des Samlandischen Unteroligozans und fiber die Ein- 

heitlichkeit der Hystrichosphaerideen. Palaeontographica, Stuttgart, (A) 105: 49-95, pis 7-12. 

1958. Mikroplankton aus dem norddeutschen Apt nebst einigen Bemerkungen fiber fossile 

Dinoflagellaten. Neues Jb. Geol. Paldont. Abh., Stuttgart, 106 (3): 383-422, pis 21-27. 

1963a. Zur Membranilarnax-Frage. Neues Jb. Geol. Paldont. Mh., Stuttgart, 1963 (2): 98-103. 

1963b. Cordosphaeridium n.g. ex Hystrichosphaeridium, Hystrichosphaeridea. Neues Jb. Geol. 

Paldont. Abh., Stuttgart, 118: 260-265, pi. 29. 

1969. Kritische Bemerkungen und Richtigstellungen im Gebiet der fossilen Dinoflagellaten 

und Acritarchen. Neues J b. Geol. Paldont. Abh., Stuttgart, 134: 101-116, pis 18-19. 

& Cookson, I. C. i960. Microplankton from Australian Lower Cretaceous sediments. Proc. R. 

Soc. Vict., Melbourne, 72: 1-11, pis 1-3. 
& Gocht, H. i960. NeueNamen ffir einige Hystrichospharen der Bernsteinformation Ostpreus- 
sens. Neues Jb. Geol. Paldont. Mh., Stuttgart, i960 (11): 511-518. 
& Kjellstrom, G. 1971. Katalog der fossilen Dinoflagellaten , Hystrichospharen und verwandten 

Mikrofossilien. Band II. Dinoflagellaten. 1130 pp., pis 1-6. 2. Ergdnzungslieferung, 215 pp. 

Stuttgart. 
Erkmen, U. & Sarjeant, W. A. S. 1980. Dinoflagellate cysts, acritarchs and tasminitids from the 

uppermost Callovian of England and Scotland: with a reconsideration of the "Xanthidium 

pilosum" problem. Geobios, Lyon, 13: 45-99, pis 1-8. 
Evitt, W. R. 1961. Observations on the morphology of fossil dinoflagellates. Micropaleontology , 

New York, 7: 385-420, pis 1-9. 

1963. A discussion and proposals concerning fossil dinoflagellates, hystrichospheres, and 

acritarchs, I. Proc. natn. Acad. Sci. U.S.A., Washington, 49: 158-164. 

1967. Dinoflagellate studies II. The archeopyle. Stanf. Univ. Pubis, (Geol. Sci.) 10 (3): 1-82, 

pis 1-11. 



50 SECOND APPENDIX TO 

Evitt, W. R., Clarke, R. F. A. & Verdier, J. -P. 1967. Dinoflagellate studies III. Dinogymnium 

acuminatum n.gen., n.sp. (Maastrichtian) and other fossils formerly referable to Gymnodinium 

Stein. Stanf. Univ. Pubis, (Geol. Sci.) 10 (4): 1-27, pis 1-3. 
Fensome, R. A. 1981. The Jurassic dinoflagellate genera Wanaea and Energlynia: their morphology 

and evolution. Neues Jb. Geol. Palaont. Abh., Stuttgart, 161: 47-61. 
Firtion, F. 1952. LeCenomanien inferieurdu Nouvion-en-Thierache: examen micropaleontologique. 

Annls Soc. geol. N., Lille, 72: 150-164, pis 8-10. 
Fisher, M. J. & van Helden, B. G. T. 1979. Some observations on the fossil dinocyst genus 

Rhaetogonyaulax Sarjeant, 1966. Palynol., Dallas, 3: 265-276, pis 1-4. 
Gerlach , E. 196 1 . Mikrofossilien aus dem Oligozan und Miozan Nordwestdeutschlands, unter beson- 

derer Beriicksichtigungder Hystrichospharen und Dinoflagellaten. Neues Jb. Geol. Palaont. Abh., 

Stuttgart, 112: 143-228, pis 25-29. 
Gocht, H. 1955. Rhombodinium und Dracodinium, zwei neue Dinoflagellaten-Gattungen aus dem 

norddeutschen Tertiar. Neues Jb. Geol. Palaont. Mh., Stuttgart, 1955 (2): 84-92, 5 figs. 
— 1959. Mikroplankton aus dem nordwestdeutschen Neokom II. Palaeont. Z., Berlin, 33: 50-89, 

pis 3-8. 
1968. Zur Morphologie und Ontogenie von Thalassiphora (Dinoflagellata). Palaeontographica , 

Stuttgart, (A) 129: 149-156, pis 25-27. 

1969. Formengemeinschaften Alttertiaren Mikroplanktons aus Bohrproben des Erdolfeldes 

Meckelfeld bei Hamburg. Palaeontographica, Stuttgart, (B) 126: 1-100, pis 1-11. 

1970. Dinoflagellaten-Zysten aus dem Bathonium des Erdolfeldes Aldorf (NW-Deutschland). 

Palaeontographica , Stuttgart, (B) 129: 125-165, pis 26-35. 

1976- Hystrichosphaeropsis quasicribrata (O. Wetzel), ein Dinoflagellat aus dem Maastricht 

Nordeuropas. Neues Jb. Geol. Palaont. Mh., Stuttgart, 1976 (6): 321-336. 

Graham, H. W. 1942. Studies in the morphology, taxonomy and ecology of the Peridiniales. Pubis 

Carnegie Inst., Washington, 542. 129 pp. 
Harland, R. 1979. The Wetzeliella (Apectodinium) homomorphum plexus from the Palaeogene/ 

earliest Eocene of north-west Europe. Proc. IV int. Palynol. Con/., Lucknow (1976-77), 2: 59- 

70, pis 1-2. 
, Morbey, S. J. & Sarjeant, W. A. S. 1975. A revision of the Triassic to lowest Jurassic 

dinoflagellate Rhaetogonyaulax. Palaeontology, London, 18: 847-864, pis 100-104. 
Ioannides, N. S., Stavrinos, G. N. & Downie, C. 1976. Kimmeridgian microplankton from 

Gavel's Hard, Dorset, England. M icropaleontology , New York, 22: 443-478. 
"Jiabo" ( = Song Zhichen et al.) 1978. [On the Paleogene dinoflagellates and acritarchs from the coastal 

region of Bohai.} 190 pp., 49 pis. Nanking Instutute of Geology and Palaeontology, Academia 

Sinica. [In Chinese: English abstract]. 
Klement, K. W. i960. Dinoflagellaten und Hystrichosphaerideen aus dem unteren und mittleren 

Malm Siidwestdeutschlands. Palaeontographica, Stuttgart, (A) 1 14: 1-104, pis 1-10. 
Klumpp, B. 1953. Beitrag zur Kenntnis der Mikrofossilien des mittleren und oberen Eozan. Palaeon- 
tographica, Stuttgart, (A) 103: 377-406, pis 16-20. 
Kofoid, C. A. 191 1. Dinoflagellata of the San Diego region, IV. The genus Gonyaulax, with notes on 

its skeletal morphology and a discussion of its generic and specific characters. Univ. Calif. 

Pubis Zool., Berkeley, 8: 187-286, pis 9-17. 
Kufferath, H. 1950. Recherches sur le plancton de la mer flamande (mer du Nord meridionale). I. 

Quelques flagelles, protistes et "caetera". Bull. Inst. r. Sci. nat. Belg., Brussels, 26: 1-43. 
Leffingwell, H. A. & Morgan, R. P. 1977. Restudy and comparison of the dinoflagellate cyst 

genus Carpodinium to that of Prionodinium n. gen. /. Paleont., Tulsa, 51: 288-302, pis 1-4. 
Lejeune-Carpentier, M. 1938. L'etude microscopique des silex. Areoligera: nouveau genre 

d'Hystrichosphaeridee (Sixieme note). Annls Soc. geol. Belg., Liege, 62: B163-B174. 

— 1939. L'etude microscopique des silex. Un nouveau Peridinien cretacique: Gonyaulax wetzeli 
(Septieme note). Annls Soc. geol. Belg., Liege, 62: B525-B529. 

— 1940. L'etude microscopique des silex. Systematiqueet morphologie des "Tubiferes" (Huitieme 
note). Annls Soc. geol. Belg., Liege, 63: B216-B236. 

— 1946. L'etude microscopique des silex. Especes nouvelles ou douteuses de Gonyaulax (Douzieme 
note). Annls Soc. geol. Belg., Liege, 69: B187-B197. 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 51 

Lejeune-Carpentier, M. & Sarjeant, W. A. S. 1981. Restudy of some larger dinoflagellate cysts 

and an acritarch from the Upper Cretaceous of Belgium and Germany. Annls Soc. geol. Belg., 

Liege, 104: 1-39, pis 1-5, text-figs 1-12. 
Lentin, J. K. & Williams, G. L. 1973. Fossil dinoflagellates: index to genera and species. Geol. 

Surv. Pap. Can., Ottawa, 73-42: 1-176. 
1975. Fossil dinoflagellates: index to genera and species. Supplement 1. Can. J. Bot., 

Ottawa, 53: 2147-2157. 

1976. A monograph of fossil peridinioid dinoflagellate cysts. Rep. Bedford Inst. Oceanogr. 



BI-R-75-16: 1-237 

1977. Fossil dinoflagellates: index to genera and species, 1977 edition. Rep. Bedford Inst. 



Oceanogr. BI-R-77-8: 1-209. 

1981. Fossil dinoflagellates: index to genera and species, 1981 edition. Rep. Bedford Inst. 



Oceanogr. BI-R-81-12: 1-345. 
Loeblich, A. R. jr & Loeblich, A. R. in 1966. Index to the genera, subgenera, and sections of the 

Pyrrhophyta. Stud. trop. Oceanogr., Miami, 3. x+94 pp., plate. 
1968. Index to the genera, subgenera, and sections of the Pyrrhophyta, II. J. Paleont., 

Menasha, 42: 210-213. 
Macro, S. 1957. Lower Miocene pollen flora from the valley of KtodnicanearGliwice (Upper Silesia). 

Pr. wroct. Tow. nauk., (ser. B) 88: 1-313, pis 1-80. 
Maier, D. 1959. Planktonuntersuchungen in tertiaren und quartaren marinen Sedimenten. Ein 

Beitrag zur Systematik, Stratigraphie und Okologie der Coccolithophorideen, Dinoflagellaten 

und Hystrichsophaerideen vom Oligozan bis zum Pleistozan. Neues Jb. Geol. Paldont. Abh., 

Stuttgart, 107: 278-340, pis 27-33. 
Mantell, G. A. 1850. A pictorial atlas of fossil remains, consisting of coloured illustrations selected 

from Parkinson's "Organic Remains of a Former World", and Artis's "Antediluvian Phytology". 

xii + 207 pp. , 74 pis. London. 
1857. Spiniferites. In: The wonders of geology; or a familiar exposition of geological phenomena 

(7th edn): 311-313, text-figs 58-59. London. 
Manum, S. & Cookson, I. C. 1964. Cretaceous microplankton in a sample from Graham Island, 

Arctic Canada, collected during the second "Fram"-Expedition (1898-1902). With notes on 

microplankton from the Hassel Formation, Ellef Ringnes Island. Skr. norske Vidensk-Akad. 

mat.-nat. Kl., Oslo, (n.s.) 17: 1-35, pis 1-7. 
Maslov, V. P. 1956. Isokopaemye isvestkovye vodorosly SSSR. Trudy Inst. geol. Nauk Mosk., 160. 

297 pp., 86 pis. 
May, F. E. 1980. Dinoflagellate cysts of the Gymnodiniaceae, Peridiniaceae, and Gonyaulacaceae 

from the Upper Cretaceous Monmouth Group, Atlantic Highlands, New Jersey. Palaeontogra- 

phica, Stuttgart, (B) 172: 10-116, pis 1-23. 
McIntyre, D. J. & Brideaux, W. W. 1980. Valanginian miospore and microplankton assemblages 

from the northern Richardson Mountains, District of Mackenzie, Canada. Bull. geol. Surv. Can., 

Ottawa, 320: 1-57, pis 1-12. 
McLean, D. M. 1973. Emendation and transfer of Eisenackia (Pyrrhophyta) from the Microdini- 

aceae to the Gonyaulacaceae. Geol. For. Stockh. Forh., 95: 261-265. 
Merrill, J. A. 1895. Fossil sponges of the flint nodules in the Lower Cretaceous of Texas. Bull. 

Mus. comp. Zool. Harv., Cambridge, Mass., 28 (1): 1-26, pi. 1. 
Morgenroth, P. 1966. Mikrofossilien und Konkretionen des nordwesteuropaischen Untereozans. 

Palaeontographica, Stuttgart, (B) 119: 1-53, pis 1-11. 
Neale, J. W. & Sarjeant, W. A. S. 1962. Microplankton from the Speeton Clay of Yorkshire. Geol. 

Mag., London, 99: 439-458, pis 19-20. 
Norris, G. 1965. Archeopyle structures in Upper Jurassic dinoflagellates from southern England. 

N.Z.Jl Geol. Geophys., Wellington, 8: 792-806. 
& Sarjeant, W. A. S. 1965. A descriptive index of genera of fossil Dinophyceae and Acritarcha. 

Palaeont. Bull. Wellington, 40. 72 pp. 
Pastiels, A. 1948. Contributions a l'etude des microfossiles de l'Eocene beige. Mem. Mus. r. Hist. 

nat. Belg., Brussels, 109: 1-77, pis 1-6. 



52 SECOND APPENDIX TO 

Philippot, A. 1949. Contributions a la paleontologiedessilexcretaces. Trois nouveaux microfossiles. 

Bull. Soc. sci. Bretagne, Rennes, 24: 55-58, 3 figs. 
Pocock, S. A. J. 1962. Microfloral analysis and age determination of strata at the Jurassic- 
Cretaceous boundary in the western Canada plains. Palaeontographica, Stuttgart, (B) III: 1- 

95.PISI-15. 
Pritchard, A. 1841. A history of Infusoria, living and fossil, arranged according to "Die Infusoriens- 

thierchen" of C . G. Ehrenberg. 439 pp., 12 pis. London. 
Reade, J. B. 1839. On some new organic remains in the Flint of Chalk. Ann. nat. Hist., London, 2: 

191-198, pis 8-9. 
Reid, P. C. 1974. Gonyaulacacean dinoflagellate cysts from the British Isles. Nova Hedwigia, 

Weinheim, 25: 579-637, pis 1-4. 
Rossignol, M. 1962. Analyse pollinique de sediments marins Quaternaires en Israel II. Sediments 

Pleistocenes. Pollen Spores, Paris, 4: 121-148, pis 1-2. 

1964. Hystrichospheres du Quaternaire en Mediterranee orientale, dans les sediments 

Pleistocenes et les boues marines actuelles. Rev. Micropaleont., Paris, 7: 83-99, P^ s I_ 3- 

Sarjeant, W. A. S. 1959. Microplankton from the Cornbrash of Yorkshire. Geol. Mag., London, 96: 

329-346, pi. 13. 
i960. New hystrichospheres from the Upper Jurassic of Dorset. Geol. Mag., London, 97: 137- 

144, pi. 6. 
1961. Microplankton from the Kellaways Rocks and Oxford Clay of Yorkshire. Palaeontology, 

London, 4: 90-118, pis 13-15. 
1962a. Upper Jurassic microplankton from Dorset, England. Micropaleontology, New York, 8: 

255-258, pis 1-2. 

1962b. Microplankton from the Ampthill Clay of Melton, South Yorkshire. Palaeontology, 

London, 5: 478-497, pis 69-70. 

1963. Two new Jurassic species of Gonyaulax (Dinophyceae). Rev. Micropaleont., Paris, 6: 85- 

88, pi. 1. 
1964. New name and diagnosis for an Upper Jurassic species of Gonyaulacysta (Dinophyceae). 

Palaeontology, London, 7: 472-473. 
1966a. The supposed "sponge spicules" of Merrill, 1895, from the Lower Cretaceous (Albian) 

of Texas. Breviora, Cambridge, Mass., 242: 1-15, pi. 1. 

1966b. Dinoflagellate cysts with Gonyaulax-type tabulation. In Davey, R. J., Downie, C, 

Sarjeant, W. A. S. & Williams, G. L. Studies on Mesozoic and Cainozoic dinoflagellate cysts. 
Bull. Br. Mus. nat. Hist., London, (Geol. Suppl.) 3: 107-156. 

1966c. Further dinoflagellate cysts from the Speeton Clay. In Davey, R. J., Downie, C, 

Sarjeant, W. A. S. & Williams, G. L. Studies on Mesozoic and Cainozoic dinoflagellate cysts. 
Bull. Br. Mus. nat. Hist., London, (Geol. Suppl.) 3: 199-214. 

1967. The genus Palaeoperidinium Deflandre (Dinophyceae). Grana palynol., Stockholm, 7: 

243-258. 

1968. Microplankton from the Upper Callovian and Lower Oxfordian of Normandy. Rev. 

Micropaleont., Paris, 10: 221-242, pis 1-3. 

1969. Taxonomic changes. In Davey, R. J., Downie, C, Sarjeant, W. A. S. & Williams, G. L. 

Appendix to "Studies on Mesozoic and Cainozoic dinoflagellate cysts." Bull. Br. Mus. nat. 

Hist., London, (Geol. Suppl.) 3 Appendix: 7-15. 
1970. The genus Spiniferites Mantell 1850 (Dinophyceae). Grana, Stockholm, 10: 74-78. 

1975. Jurassic dinoflagellate cysts with epitractal archeopyles. A reconsideration. Grana, 

Stockholm, 14: 49-56, pis 1-3. 

1976. English Jurassic dinoflagellate cysts and acritarchs: a re-examination of some type and 

figured specimens. Geosci. Man, Baton Rouge, 15: 1-24, pis 1-7. 
— 1980. A restudy of some dinoflagellate cyst holotypes in the University of Kiel collections. I. 
The Jurassic holotypes of Walter Wetzel (1966a, b). Meyniana, Kiel, 32: 113-128, pis 1-3. 
— 1981. A restudy of some dinoflagellate cyst holotypes in the University of Kiel collections. II. 
The Eocene holotypes of Barbara Klumpp (1953); with a revision of the genus Cordosphaeri- 
dium Eisenack (1953). Meyniana, Kiel, 33: 97-132. 



DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 53 

Sarjeant, W. A. S. 1982a. The dinoflagellate cysts of the Gonyaulacysta group: a morphological 

and taxonomic restudy. Contr. Ser. am. Ass. stratigr. Palynol., Dallas, 9. ii + 81 pp., 12 pis. 

1982b. Dinoflagellate cyst terminology: a discussion and proposals. Can. J . Bot., Ottawa, 60 

(6): 922-945. 

1983a. A restudy of some dinoflagellate cyst holotypes in the University of Kiel collections. 

III. The Oligocene and Miocene holotypes of Dorothea Maier (1959). Meyniana, Kiel. (In 

Press). 

1983b. A restudy of some dinoflagellate cysts and an acritarch from the Malm (Upper Jurassic) 

of southwest Germany. Palaeontographica, Stuttgart, (B). (In Press). 
& Stover, L. E. 1978. Cyclonephelium and Tenua: a problem in dinoflagellate cyst taxonomy. 

Grana, Stockholm, 17: 47-54. 
Staplin, F. L., Jansonius, J. & Pocock, S. A. J. 1965. Evaluation of some acritarchous hystri- 

chosphere genera. Neues Jb.Geol.Paldont. Abh., Stuttgart, 123: 167-201, pis 18-20. 
Stover, L. E. 1975. Observations on some Australian Eocene dinoflagellates. Geosci. Man, Baton 

Rouge, 11:35-45, pis 1-3. 
& Evitt, W. R. 1978. Analyses of pre- Pleistocene organic-walled dinoflagellates. Stan/. Univ. 

Pubis, (Geol. Sci.) 15: 1-300. 
Takahashi, K. 1964. Microplankton from the Asagai Formation in the Joban Coalfield. Trans. 

Proc. palaeont. Soc. Japan, Tokyo, (N.S) 54: 201-214, pis 30-33. 
Tasch, P. 1964. In Tasch, P., McClure, K. & Oftedahl, O., Biostratigraphy and taxonomy of a 

hystrichosphere-dinoflagellate assemblage from the Cretaceous of Kansas. Micropaleontology , 

New York, 10: 189-206, pis 1-3. 
Valensi, L. 1947. Note preliminaire a une etude des microfossils des silex jurassiques de le region de 

Poitiers. C. r. hebd. Seanc. Acad. Sci., Paris, 225: 816-818. 
1948. Sur quelques microorganismes planctoniques des silex du Jurassique moyen du Poitou 

et de Normandie. Bull. Soc. geol. Fr., Paris, (5) 18: 537-550. 
1953 Microfossiles des silex du Jurassique moyen. Remarques petrographiques. Mem. Soc. geol. 

Fr., Paris, 68. 100 pp., 16 pis. 
1955- Etude micropaleontologique des silex du Magdalenien de Saint-Amand (Cher). Bull. Soc. 

prehist.fr., Paris, 52: 584-596, pis 1-5. 
Vozzhennikova, T. F. 1967. I skopaemye perdinei yurskikh, melovykh i paleogenovykh otlozheniy 

SSSR. 347 pp., 121 pis. Moscow (Nauka). [In Russian: Engl, transl. Fossilized peridinid algae 

in the Jurassic, Cretaceous and Palaeogene deposits of the USSR. 1971. 453 pp., 121 pis. 

National Lending Library for Science and Technology, Boston Spa]. 
Wall, D. 1967. Fossil microplankton in deep-sea cores from the Caribbean Sea. Palaeontology, 

London, 10: 95-123, pis 14-16. 
Warren, J. S. 1973. Form and variation of the dinoflagellate Sirmiodinium grossi Alberti, from the 

Upper Jurassic and Lower Cretaceous of California. /. Paleont., Tulsa, 47: 101-114, pis 1-3. 
Weiler, H. 1956. Uber einen Fund von Dinoflagellaten, Coccolithophoriden und Hystrichosphaeri- 

deen im Tertiar des Rheintales. Neues Jb. Geol. Paldont. Mh., Stuttgart, 104: 129-147, pis 

11-13. 
Wetzel, O. 1933. Die in organischer Substanz erhaltenen Mikrofossilien des Baltischen Kreide- 

Feuersteins. Palaeontographica, Stuttgart, 77: 141-188; (A) 78: i-no, pis 1-7. 
Wetzel, W. 1952. Beitrag zur Kenntnis des dan-zeitlichen Meeresplanktons. Geol. Jb., Hannover, 

66: 391-419, pi. A. 
1967. Mikroorganismen aus Jurassischen und Kretazischen Saurier-Gewollan. Z. dt. geol. Ges., 

Berlin, 116: 867-874, pis 1-3. 
White, H. H. 1842. On fossil Xanthidia. Microsc. J. & struct. Rec, London, 2: 35-40, pi. 4. 
Wiggins, V. D. 1972. Two new Lower Cretaceous dinoflagellate genera from southern Alaska 

(U.S.A.). Rev. Palaeobot. Palynol., Amsterdam, 14: 297-308, pis 1-3. 
1975- The dinoflagellate Family Pareodiniaceae: a discussion. Geosci. Man, Baton Rouge, n: 

95-115, pis 1-5. 
Wilkinson, S. J. 1849. Observations on Xanthidium, both Fossil and Recent. Trans, microsc. Soc. 

Lond., 2: 89-92, pi. 13. 



54 SECOND APPENDIX 

Williams, G. L. & Downie, C. 1966a. The genus Hystrichokolpoma. In Davey, R. J., Downie, C, 

Sarjeant, W. A. S. & Williams, G. L. Studies on Mesozoic and Cainozoic dinoflagellate cysts. 

Bull. Br. Mus. nat. Hist., London, (Geol. Suppl.) 3: 176-181. 
1966b. Wetzeliella from the London Clay. In Davey, R. J., Downie, C, Sarjeant, W. A. S. 

& Williams, G. L. Studies on Mesozoic and Cainozoic dinoflagellate cysts. Bull. Br. Mus. nat. 

Hist., London, (Geol. Suppl.) 3: 182-198. 

1966c. Further dinoflagellate cysts from the London Clay. In Davey, R. J., Downie, C. 



Sarjeant, W. A. S. & Williams, G. L. Studies on Mesozoic and Cainozoic dinoflagellate cysts. 
Bull. Br. Mus. nat. Hist., London, (Geol. Suppl.) 3: 215-235. 

-1969. Generic re-allocations. In Davey, R. J., Downie, C, Sarjeant, W. A. S. & Williams, 



G. L. Appendix to "Studies on Mesozoic and Cainozoic dinoflagellate cysts." Bull. Br. Mus. 
nat. Hist., London, (Geol. Suppl.) 3 Appendix: 17. 

-, Sarjeant, W. A. S. & Kidson, E. J. 1978. A glossary of the terminology applied to dinofla- 
gellate amphiesmae, and cysts and acritarchs. 1978 Edition. Contr. Ser. am. Ass. stratigr. 
Palynol., Dallas, 2A. 121 pp., 15 pis. 



Accepted for publication 15 December 1982 



L. A. Krukewich, B.Sc, 

W. A. S. Sarjeant, B.Sc, Ph.D., F.G.S., F.L.S. 

Dept. of Geological Sciences 

University of Saskatchewan 

Saskatoon 

Canada